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Large equity awards granted to Aura Biosciences (AURA) director Natalie Holles

Filing Impact
(Neutral)
Filing Sentiment
(Neutral)
Form Type
4

Rhea-AI Filing Summary

Aura Biosciences, Inc. reported that director and officer Natalie C. Holles received significant equity awards in the form of restricted stock units, performance RSUs and stock options. She was granted 553,844 RSUs as Inducement Awards, each representing one share of common stock, vesting in four equal annual installments beginning on April 15, 2027, subject to continued service.

She also received 600,118 performance RSUs as Inducement Awards that require both time-based vesting and achievement of a performance condition. These PRSUs vest in four equal annual installments commencing on April 15, 2027 and expire on the earlier of the sixth anniversary of the grant date or the date her service with the company ends, with any unvested awards then forfeited.

In addition, Holles was granted a stock option covering 2,169,103 shares of common stock at an exercise price of $7.3900 per share, expiring on May 19, 2036. Twenty-five percent of the option vests on April 30, 2027, with the remainder vesting pro rata in 36 monthly installments, subject to continued service. Following these grants, she directly owns 1,153,962 shares of common stock.

Positive

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Insider Holles Natalie C.
Role See Remarks
Type Security Shares Price Value
Grant/Award Stock Option (Right to Buy) 2,169,103 $0.00 --
Grant/Award Common Stock 600,118 $0.00 --
Grant/Award Common Stock 553,844 $0.00 --
Holdings After Transaction: Stock Option (Right to Buy) — 2,169,103 shares (Direct, null); Common Stock — 600,118 shares (Direct, null)
Footnotes (1)
  1. These shares were acquired pursuant to a restricted stock unit ("RSU") award as Inducement Awards. Each RSU represents the right to receive one share of the Issuer's common stock. These RSUs vest in four substantially equal annual installments beginning on April 15, 2027, subject to the Reporting Person's continued service as of each such vesting date. These shares were acquired pursuant to a performance RSU ("PRSU") award as Inducement Awards. The PRSUs are subject to both time-based vesting and the achievement of a performance condition, both of which must be satisfied before the PRSUs will be deemed vested. The PRSUs shall vest in four substantially equal annual installments commencing on April 15, 2027, subject to Ms. Holles' continued service as of each such time-based vesting date and the satisfaction of the performance condition. The expiration date of the PRSUs is the earlier of (i) the sixth (6th) anniversary of the grant date and (ii) the date Ms. Holles no longer has a service relationship with the Issuer. Any such PRSUs that have not vested on or prior to such expiration date shall be forfeited for no consideration. The shares underlying this option vest as follows: 25% of the shares vest on April 30, 2027 with the remainder vesting thereafter pro-rata in 36 monthly installments, subject to the Reporting Person's continued service as of each such vesting date.
RSU grant 553,844 shares RSU Inducement Award vesting annually from April 15, 2027
Performance RSU grant 600,118 shares PRSU Inducement Award with time and performance conditions
Stock option size 2,169,103 shares Option to buy common stock granted to Holles
Option exercise price $7.3900 per share Exercise price of stock option expiring May 19, 2036
Option expiration May 19, 2036 Expiration date of stock option grant
Post-award holdings 1,153,962 shares Common stock directly owned after non-derivative acquisitions
restricted stock unit ("RSU") financial
"These shares were acquired pursuant to a restricted stock unit ("RSU") award as Inducement Awards."
performance RSU ("PRSU") financial
"These shares were acquired pursuant to a performance RSU ("PRSU") award as Inducement Awards."
Inducement Awards financial
"These shares were acquired pursuant to a restricted stock unit ("RSU") award as Inducement Awards."
Inducement awards are special bonuses given to new employees to encourage them to join a company, often in the form of stock or money. They matter because they can motivate talented people to choose one company over another and help align their success with the company's growth. Think of it like a signing bonus to seal the deal.
service relationship financial
"the date Ms. Holles no longer has a service relationship with the Issuer."
Stock Option (Right to Buy) financial
"Stock Option (Right to Buy)"
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Learn about SEC filing dates
SEC Form 4
FORM 4UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934
or Section 30(h) of the Investment Company Act of 1940
OMB APPROVAL
OMB Number:3235-0287
Estimated average burden
hours per response:0.5
Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).
Check this box to indicate that a transaction was made pursuant to a contract, instruction or written plan for the purchase or sale of equity securities of the issuer that is intended to satisfy the affirmative defense conditions of Rule 10b5-1(c). See Instruction 10.
1. Name and Address of Reporting Person*
Holles Natalie C.

(Last)(First)(Middle)
C/O AURA BIOSCIENCES, INC.
80 GUEST STREET

(Street)
BOSTON MASSACHUSETTS 02135

(City)(State)(Zip)

UNITED STATES

(Country)
2. Issuer Name and Ticker or Trading Symbol
Aura Biosciences, Inc. [ AURA ]
5. Relationship of Reporting Person(s) to Issuer
(Check all applicable)
XDirector10% Owner
XOfficer (give title below)Other (specify below)
See Remarks
2a. Foreign Trading Symbol
3. Date of Earliest Transaction (Month/Day/Year)
05/19/2026
6. Individual or Joint/Group Filing (Check Applicable Line)
XForm filed by One Reporting Person
Form filed by More than One Reporting Person
4. If Amendment, Date of Original Filed (Month/Day/Year)

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1. Title of Security (Instr. 3) 2. Transaction Date (Month/Day/Year)2A. Deemed Execution Date, if any (Month/Day/Year)3. Transaction Code (Instr. 8) 4. Securities Acquired (A) or Disposed Of (D) (Instr. 3, 4 and 5) 5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4) 6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4) 7. Nature of Indirect Beneficial Ownership (Instr. 4)
CodeVAmount(A) or (D)Price
Common Stock05/19/2026A600,118(1)A$0600,118D
Common Stock05/19/2026A553,844(2)A$01,153,962D
Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned
(e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security (Instr. 3) 2. Conversion or Exercise Price of Derivative Security 3. Transaction Date (Month/Day/Year)3A. Deemed Execution Date, if any (Month/Day/Year)4. Transaction Code (Instr. 8) 5. Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5) 6. Date Exercisable and Expiration Date (Month/Day/Year)7. Title and Amount of Securities Underlying Derivative Security (Instr. 3 and 4) 8. Price of Derivative Security (Instr. 5) 9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4) 10. Ownership Form: Direct (D) or Indirect (I) (Instr. 4) 11. Nature of Indirect Beneficial Ownership (Instr. 4)
CodeV(A)(D)Date ExercisableExpiration DateTitleAmount or Number of Shares
Stock Option (Right to Buy)$7.3905/19/2026A2,169,103 (3)05/19/2036Common Stock2,169,103$02,169,103D
Explanation of Responses:
1. These shares were acquired pursuant to a restricted stock unit ("RSU") award as Inducement Awards. Each RSU represents the right to receive one share of the Issuer's common stock. These RSUs vest in four substantially equal annual installments beginning on April 15, 2027, subject to the Reporting Person's continued service as of each such vesting date.
2. These shares were acquired pursuant to a performance RSU ("PRSU") award as Inducement Awards. The PRSUs are subject to both time-based vesting and the achievement of a performance condition, both of which must be satisfied before the PRSUs will be deemed vested. The PRSUs shall vest in four substantially equal annual installments commencing on April 15, 2027, subject to Ms. Holles' continued service as of each such time-based vesting date and the satisfaction of the performance condition. The expiration date of the PRSUs is the earlier of (i) the sixth (6th) anniversary of the grant date and (ii) the date Ms. Holles no longer has a service relationship with the Issuer. Any such PRSUs that have not vested on or prior to such expiration date shall be forfeited for no consideration.
3. The shares underlying this option vest as follows: 25% of the shares vest on April 30, 2027 with the remainder vesting thereafter pro-rata in 36 monthly installments, subject to the Reporting Person's continued service as of each such vesting date.
Remarks:
Chief Executive Officer and President
/s/ Conor Kilroy, as Attorney-in-Fact05/19/2026
** Signature of Reporting PersonDate
Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
* If the form is filed by more than one reporting person, see Instruction 4 (b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
Note: File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB Number.
* Form 4: SEC 1474 (03-26)

FAQ

What equity awards did Natalie C. Holles receive from Aura Biosciences (AURA)?

Natalie C. Holles received RSUs, performance RSUs, and stock options from Aura Biosciences. She was granted 553,844 RSUs, 600,118 performance RSUs, and a stock option for 2,169,103 shares, all as Inducement Awards tied to multi-year vesting schedules and continued service.

How do the RSUs granted to Natalie C. Holles at Aura Biosciences vest?

The 553,844 RSUs granted to Holles vest over four years starting April 15, 2027. They vest in four substantially equal annual installments, and each RSU converts into one share of Aura Biosciences common stock, assuming she continues serving through each vesting date.

What conditions apply to Natalie C. Holles’ performance RSUs at Aura Biosciences (AURA)?

The 600,118 performance RSUs require both time-based vesting and meeting a performance condition. They vest in four equal annual installments beginning April 15, 2027, only if Holles remains in service and the specified performance condition is satisfied before the awards are deemed vested.

What are the key terms of Natalie C. Holles’ stock option grant from Aura Biosciences?

Holles received a stock option to buy 2,169,103 Aura Biosciences shares at $7.39. Twenty-five percent vests on April 30, 2027, with the remaining portion vesting pro rata over 36 monthly installments, and the option expires on May 19, 2036, subject to continued service.

When do Natalie C. Holles’ performance RSUs at Aura Biosciences expire if unvested?

The performance RSUs expire at the earlier of six years from grant or her service ending. Any performance RSUs that have not vested by the sixth anniversary or by the time she no longer has a service relationship with Aura Biosciences are forfeited for no consideration.

How many Aura Biosciences shares does Natalie C. Holles hold after these awards?

After the reported awards, Holles directly owns 1,153,962 shares of Aura Biosciences common stock. This total reflects the non-derivative holdings shown following the grants of restricted stock units and performance RSUs disclosed in the Form 4 filing.