CDXS insider filing: director sells 1,413 shares to cover taxes

Rhea-AI Filing Summary

Codexis (CDXS) reported a Form 4 for director Raymond De Vre. On 11/11/2025, he sold 1,413 shares of common stock at $1.74 per share, coded “S.” The filing notes the sale was effected to cover tax liability from the vesting and release of restricted stock units. After the transaction, he beneficially owned 87,416 shares, which includes 72,880 restricted stock units. Ownership is reported as direct.

Insider Trade Summary

Net Seller: 1,413 shares ($2,459)

Net Sell 1 txn

Insider De Vre Raymond

Role Director

Sold 1,413 shs ($2K)

Type	Security	Shares	Price	Value
Sale	Common Stock	1,413	$1.74	$2K

Holdings After Transaction: Common Stock — 87,416 shares (Direct)

Footnotes (1)

1. The sale reported on this Form 4 was effected to cover tax liability incurred due to the vesting and release of Restricted Stock Units held by the Reporting Person. Includes 72,880 restricted stock units.

Form 4 Insider Transactions: What Each Field Means →

SEC Form 4

FORM 4

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934
or Section 30(h) of the Investment Company Act of 1940

OMB APPROVAL

OMB Number:			3235-0287
Estimated average burden
hours per response:			0.5

Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).

Check this box to indicate that a transaction was made pursuant to a contract, instruction or written plan for the purchase or sale of equity securities of the issuer that is intended to satisfy the affirmative defense conditions of Rule 10b5-1(c). See Instruction 10.

1. Name and Address of Reporting Person^*

De Vre Raymond

(Last) (First) (Middle)

200 PENOBSCOT DRIVE

(Street)

REDWOOD CITY CA 94063

(City) (State) (Zip)

2. Issuer Name and Ticker or Trading Symbol
CODEXIS, INC. [ CDXS ]

5. Relationship of Reporting Person(s) to Issuer
(Check all applicable)

X	Director		10% Owner
	Officer (give title below)		Other (specify below)

3. Date of Earliest Transaction (Month/Day/Year)
11/11/2025

4. If Amendment, Date of Original Filed (Month/Day/Year)

6. Individual or Joint/Group Filing (Check Applicable Line)

X	Form filed by One Reporting Person
	Form filed by More than One Reporting Person

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1. Title of Security (Instr. 3)	2. Transaction Date (Month/Day/Year)	2A. Deemed Execution Date, if any (Month/Day/Year)	3. Transaction Code (Instr. 8)		4. Securities Acquired (A) or Disposed Of (D) (Instr. 3, 4 and 5)			5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4)	6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	7. Nature of Indirect Beneficial Ownership (Instr. 4)
			Code	V	Amount	(A) or (D)	Price
Common Stock	11/11/2025		S(1)		1,413	D	$1.74	87,416(2)	D

Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned (e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security (Instr. 3)	2. Conversion or Exercise Price of Derivative Security	3. Transaction Date (Month/Day/Year)	3A. Deemed Execution Date, if any (Month/Day/Year)	4. Transaction Code (Instr. 8)		5. Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5)		6. Date Exercisable and Expiration Date (Month/Day/Year)		7. Title and Amount of Securities Underlying Derivative Security (Instr. 3 and 4)		8. Price of Derivative Security (Instr. 5)	9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4)	10. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	11. Nature of Indirect Beneficial Ownership (Instr. 4)
				Code	V	(A)	(D)	Date Exercisable	Expiration Date	Title	Amount or Number of Shares

Explanation of Responses:

1. The sale reported on this Form 4 was effected to cover tax liability incurred due to the vesting and release of Restricted Stock Units held by the Reporting Person.

2. Includes 72,880 restricted stock units.

	/s/ Georgia Erbez, as Attorney-in-Fact for Raymond De Vre	11/13/2025
	** Signature of Reporting Person	Date
Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
* If the form is filed by more than one reporting person, see Instruction 4 (b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
Note: File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB Number.

FAQ

What did Codexis (CDXS) disclose in this Form 4?

A director, Raymond De Vre, reported a sale of 1,413 shares of common stock.

When did the CDXS insider transaction occur?

The transaction date was 11/11/2025.

What was the sale price in the CDXS Form 4?

Shares were sold at $1.74 per share.

Why were the shares sold by the CDXS director?

The filing states the sale was to cover tax liability from the vesting and release of RSUs.

How many shares does the insider beneficially own after the sale?

Post-transaction beneficial ownership is 87,416 shares, including 72,880 RSUs.

What is the insider’s relationship to Codexis (CDXS)?

The reporting person is a Director of Codexis.

Is the ownership direct or indirect in this CDXS filing?

Ownership is reported as Direct.