Codexis director granted 66,350 stock options

CODEXIS, INC. director Esther Martinborough reported receiving a stock option grant covering 66,350 shares of common stock at an exercise price of $2.48 per share.

Accepted by SEC EDGAR 06/18/2026 - 04:32 PM EDT Learn about SEC filing dates

Rhea-AI Filing Summary

CODEXIS, INC. director Esther Martinborough reported receiving a stock option grant covering 66,350 shares of common stock at an exercise price of $2.48 per share. The option vests in full on the earlier of the first anniversary of the grant date or the next annual stockholder meeting, subject to her continued board service, and expires on June 17, 2036. Following this grant, she holds options for 66,350 shares directly.

Insider Trade Summary

Grant/Award: 66,350 shares

Grant/Award 1 txn

Insider Martinborough Esther

Role Director

Type	Security	Shares	Price	Value
Grant/Award	Stock Option (Right to Buy)	66,350	$0.00	$0.00

Holdings After Transaction: Stock Option (Right to Buy) — 66,350 options (Direct)

Holdings Before and After: Stock Option (Right to Buy), 0 to 66,350 options (Direct)

Footnotes (1)

1. F1. This option vests and becomes exercisable with respect to all of the shares subject to the award on the earlier of the first anniversary of the date of grant or the next annual stockholder meeting, subject to the director's continued service to the Company on such vesting date.

Key Figures

Option grant size: 66,350 shares Exercise price: $2.48 per share Underlying shares: 66,350 shares +2 more

5 metrics

Option grant size 66,350 shares Stock Option (Right to Buy) granted to director on June 17, 2026

Exercise price $2.48 per share Conversion or exercise price of the granted stock option

Underlying shares 66,350 shares Underlying Codexis common stock covered by this option

Post-transaction option holdings 66,350 shares Total option shares held directly following the grant

Option expiration date June 17, 2036 Expiration of the stock option grant

Key Terms

Stock Option (Right to Buy), grant/award acquisition, exercise price, Common Stock, +1 more

5 terms

Stock Option (Right to Buy) financial

"security_title: Stock Option (Right to Buy)"

grant/award acquisition financial

"transaction_action: grant/award acquisition"

exercise price financial

"conversion_or_exercise_price: 2.4800"

The exercise price is the fixed amount at which you can buy or sell an asset, like a stock, when using an options contract. It matters because it helps determine whether exercising the option will be profitable or not, depending on the current market price. Think of it as the set price you agree on today to buy or sell later.

Common Stock financial

"underlying_security_title: Common Stock"

Common stock represents ownership shares in a company, giving investors a stake in its success and a say in important decisions through voting rights. It is the most common type of stock traded on markets and can provide income through dividends, as well as potential for value growth. For investors, holding common stock means sharing in the company’s profits and risks.

vests and becomes exercisable financial

"This option vests and becomes exercisable with respect to all of the shares"

FAQ

AI-generated questions and answers. How Rhea-AI works. Not financial advice.

What did Codexis (CDXS) director Esther Martinborough report on this Form 4?

Esther Martinborough reported receiving a stock option grant for 66,350 shares of Codexis common stock. The options were awarded as director compensation and give her the right to buy shares at a fixed exercise price of $2.48 per share in the future.

How many Codexis (CDXS) shares are covered by Esther Martinborough’s new option grant?

The new grant covers options to acquire 66,350 shares of Codexis common stock. These options provide potential future ownership if exercised, giving her the right to purchase those shares at the predetermined strike price once the award becomes fully vested and exercisable.

What is the exercise price of Esther Martinborough’s Codexis (CDXS) stock options?

The options have an exercise price of $2.48 per share. This means she can choose to buy Codexis common stock at $2.48 for each optioned share once the grant vests, regardless of the market price at the time she exercises the options.

When do Esther Martinborough’s Codexis (CDXS) stock options vest?

The option vests and becomes exercisable in full on the earlier of the first anniversary of the grant date or the next annual stockholder meeting. Vesting is conditioned on her continued service as a director through the applicable vesting date under the award terms.

When do Esther Martinborough’s Codexis (CDXS) stock options expire?

The stock options expire on June 17, 2036. After that expiration date, any unexercised portion of the 66,350-share grant will lapse, and she will no longer have the right to purchase Codexis common stock under this particular option award.

How many Codexis (CDXS) option shares does Esther Martinborough hold after this transaction?

After this reported grant, she holds options for 66,350 shares of Codexis common stock directly. The Form 4 shows this entire amount as her total option position following the transaction, reflecting this single newly awarded director stock option grant.

AI-generated analysis. How Rhea-AI works. Not financial advice.

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SEC Form 4

FORM 4

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934
or Section 30(h) of the Investment Company Act of 1940

OMB APPROVAL

OMB Number:			3235-0287
Estimated average burden
hours per response:			0.5

Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).

Check this box to indicate that a transaction was made pursuant to a contract, instruction or written plan for the purchase or sale of equity securities of the issuer that is intended to satisfy the affirmative defense conditions of Rule 10b5-1(c). See Instruction 10.

1. Name and Address of Reporting Person^*

Martinborough Esther

(Last) (First) (Middle)

C/O CODEXIS, INC.

200 PENOBSCOT DRIVE

(Street)

REDWOOD CITY CALIFORNIA 94063

(City) (State) (Zip)

UNITED STATES

(Country)

2. Issuer Name and Ticker or Trading Symbol
CODEXIS, INC. [ CDXS ]

5. Relationship of Reporting Person(s) to Issuer
(Check all applicable)

X	Director		10% Owner
	Officer (give title below)		Other (specify below)

2a. Foreign Trading Symbol

3. Date of Earliest Transaction (Month/Day/Year)
06/17/2026

6. Individual or Joint/Group Filing (Check Applicable Line)

X	Form filed by One Reporting Person
	Form filed by More than One Reporting Person

4. If Amendment, Date of Original Filed (Month/Day/Year)

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1. Title of Security (Instr. 3)	2. Transaction Date (Month/Day/Year)	2A. Deemed Execution Date, if any (Month/Day/Year)	3. Transaction Code (Instr. 8)		4. Securities Acquired (A) or Disposed Of (D) (Instr. 3, 4 and 5)			5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4)	6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	7. Nature of Indirect Beneficial Ownership (Instr. 4)
			Code	V	Amount	(A) or (D)	Price

Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned (e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security (Instr. 3)	2. Conversion or Exercise Price of Derivative Security	3. Transaction Date (Month/Day/Year)	3A. Deemed Execution Date, if any (Month/Day/Year)	4. Transaction Code (Instr. 8)		5. Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5)		6. Date Exercisable and Expiration Date (Month/Day/Year)		7. Title and Amount of Securities Underlying Derivative Security (Instr. 3 and 4)		8. Price of Derivative Security (Instr. 5)	9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4)	10. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	11. Nature of Indirect Beneficial Ownership (Instr. 4)
				Code	V	(A)	(D)	Date Exercisable	Expiration Date	Title	Amount or Number of Shares
Stock Option (Right to Buy)	$2.48	06/17/2026		A		66,350		(1)	06/17/2036	Common Stock	66,350	$0	66,350	D

Explanation of Responses:

1. This option vests and becomes exercisable with respect to all of the shares subject to the award on the earlier of the first anniversary of the date of grant or the next annual stockholder meeting, subject to the director's continued service to the Company on such vesting date.

	/s/ Georgia Erbez, as Attorney-in-Fact for Esther Martinborough	06/18/2026
	** Signature of Reporting Person	Date
Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
* If the form is filed by more than one reporting person, see Instruction 4 (b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
Note: File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB Number.
* Form 4: SEC 1474 (03-26)