Codexis holder Lytton reports 5.2% ownership stake

Laurence W. Lytton reports beneficial ownership of 5,592,972 shares of Codexis, Inc. common stock, representing 5.2% of the class.

Rhea-AI Filing Summary

Laurence W. Lytton reports beneficial ownership of 5,592,972 shares of Codexis, Inc. common stock, representing 5.2% of the class. The percentage is based on 107,561,409 shares outstanding following an issuance described in a July 24, 2026 prospectus. The filing notes that as of June 30, 2026, his beneficial ownership was below 5%, but subsequent purchases increased his holdings above the 5% threshold.

Lytton reports sole voting and dispositive power over the large majority of these shares, with a smaller portion subject to shared voting and dispositive power.

Key Figures

Beneficially owned shares: 5,592,972 shares Percent of class: 5.2% Shares outstanding basis: 107,561,409 shares +1 more

4 metrics

Beneficially owned shares 5,592,972 shares Amount of Codexis common stock beneficially owned by Laurence W. Lytton

Percent of class 5.2% Portion of Codexis common stock beneficially owned by Lytton

Shares outstanding basis 107,561,409 shares Codexis common shares outstanding used to calculate Lytton’s ownership percentage

Sole voting power 5,498,680 shares Shares over which Lytton has sole power to vote or direct the vote

Key Terms

beneficial ownership, Sole Voting Power, dispositive power, Percent of class

4 terms

beneficial ownership financial

"the reporting person's beneficial ownership was less than 5%, but subsequent purchases"

Beneficial ownership means the person or entity that actually enjoys the benefits of owning shares or other assets — such as receiving dividends, voting rights, or price gains — even if the legal title is held in another name. For investors it matters because knowing who truly controls and profits from a company reveals who can influence decisions, exposes potential conflicts of interest or hidden concentration of power, and affects transparency and risk in the stock.

Sole Voting Power financial

"Sole Voting Power 5,498,680.00 6 | Shared Voting Power"

Sole voting power is the exclusive right to cast votes attached to a shareholder’s stock without needing approval from anyone else. Like holding the only remote control for a TV, it lets that holder decide corporate matters such as board members, mergers, and policy changes, making it important to investors because it concentrates control and can strongly influence a company’s strategy and the value of its shares.

dispositive power financial

"Sole Dispositive Power 5,498,680.00 8 | Shared Dispositive Power"

Dispositive power is the authority to decide the final outcome of an asset, legal claim, contract, or corporate action — in effect the power to dispose of or resolve something. For investors it matters because whoever holds that authority can determine who gets paid, who controls an asset or vote, and how risks and returns are allocated; think of it like holding the key that lets you lock in the winner or loser in a deal.

Percent of class financial

"(b) | Percent of class: 5.2%"

Percent of class is the portion of a specific category of securities—such as a company’s common shares, preferred shares, or a bond series—that takes part in or approves a corporate action (vote, consent, tender, etc.). Investors watch this number because it reveals how much support or opposition exists within that particular shareholder group; like counting how many members of a club back a proposal, it can determine whether a plan passes or how influence is distributed.

FAQ

AI-generated questions and answers. How Rhea-AI works. Not financial advice.

What stake in Codexis (CDXS) does Laurence W. Lytton report in this Schedule 13G/A?

Laurence W. Lytton reports beneficial ownership of 5,592,972 Codexis shares, representing 5.2% of the company’s common stock. This level of ownership makes him a significant shareholder under SEC reporting rules.

How is Laurence W. Lytton’s 5.2% Codexis (CDXS) ownership percentage calculated?

The 5.2% beneficial ownership is calculated using 107,561,409 Codexis common shares outstanding, following an issuance described in a prospectus filed on July 24, 2026. This share count serves as the denominator for the percentage.

Did Laurence W. Lytton’s Codexis (CDXS) ownership change around June 30, 2026?

Yes. The filing states that as of June 30, 2026, Lytton’s beneficial ownership was below 5%. Subsequent purchases increased his holdings, bringing his beneficial ownership above the 5% reporting threshold.

What voting and dispositive powers over Codexis (CDXS) shares does Laurence W. Lytton report?

Lytton reports sole voting and dispositive power over most of his 5,592,972 Codexis shares, and shared voting and dispositive power over a smaller portion. These distinctions indicate how he can vote and dispose of the shares.

Who is the reporting person in this Codexis (CDXS) Schedule 13G/A and what is his citizenship?

The reporting person is Laurence W. Lytton, who lists his citizenship as USA. His address is given as 467 Central Park West, New York, NY 10025, and he reports holdings in Codexis common stock.

When was this Codexis (CDXS) Schedule 13G/A signed by Laurence W. Lytton?

The Schedule 13G/A was signed by Laurence W. Lytton on August 13, 2026. The signature confirms the reported Codexis common stock ownership details as of the dates described in the filing.

AI-generated analysis. How Rhea-AI works. Not financial advice.

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Available on EDGAR 08/13/2026 - 02:09 PM Accepted by SEC EDGAR 08/13/2026 - 02:08 PM Learn about SEC filing dates

SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

SCHEDULE 13G

UNDER THE SECURITIES EXCHANGE ACT OF 1934

(Amendment No. 1)

Codexis, Inc.

(Name of Issuer)

Common Stock

(Title of Class of Securities)

192005106

(CUSIP Number)

06/30/2026

(Date of Event Which Requires Filing of this Statement)

Check the appropriate box to designate the rule pursuant to which this Schedule is filed:

Rule 13d-1(b)

Rule 13d-1(c)

Rule 13d-1(d)

schemaVersion:

SCHEDULE 13G

CUSIP Number(s): 192005106

1

Names of Reporting Persons

LYTTON LAURENCE W

2

Check the appropriate box if a member of a Group (see instructions)

(a)
(b)

3

Sec Use Only

4

Citizenship or Place of Organization

UNITED STATES

Number of Shares Beneficially Owned by Each Reporting Person With:

5	Sole Voting Power 5,498,680.00
6	Shared Voting Power 94,292.00
7	Sole Dispositive Power 5,498,680.00
8	Shared Dispositive Power 94,292.00

9

Aggregate Amount Beneficially Owned by Each Reporting Person

5,592,972.00

10

Check box if the aggregate amount in row (9) excludes certain shares (See Instructions)

11

Percent of class represented by amount in row (9)

5.2 %

12

Type of Reporting Person (See Instructions)

IN

Comment for Type of Reporting Person: The reporting persons are reporting their holdings as of the date of this filing. As of June 30, 2026, the reporting person's beneficial ownership was less than 5%, but subsequent purchases caused his beneficial ownership to increase. Percentage calculated based on 107,561,409 shares of Common Stock outstanding following the issuance of shares by the Issuer described in the Prospectus filed by the Issuer on July 24, 2026.

SCHEDULE 13G

Item 1.
(a)	Name of issuer: Codexis, Inc.
(b)	Address of issuer's principal executive offices: 200 Penobscot Drive, Redwood City, CA 94063
Item 2.
(a)	Name of person filing: Laurence W. Lytton
(b)	Address or principal business office or, if none, residence: 467 Central Park West New York, NY 10025
(c)	Citizenship: USA
(d)	Title of class of securities: Common Stock
(e)	CUSIP No.: 192005106
Item 3.	If this statement is filed pursuant to §§ 240.13d-1(b) or 240.13d-2(b) or (c), check whether the person filing is a:
(a)	Broker or dealer registered under section 15 of the Act (15 U.S.C. 78o);
(b)	Bank as defined in section 3(a)(6) of the Act (15 U.S.C. 78c);
(c)	Insurance company as defined in section 3(a)(19) of the Act (15 U.S.C. 78c);
(d)	Investment company registered under section 8 of the Investment Company Act of 1940 (15 U.S.C. 80a-8);
(e)	An investment adviser in accordance with § 240.13d-1(b)(1)(ii)(E);
(f)	An employee benefit plan or endowment fund in accordance with § 240.13d-1(b)(1)(ii)(F);
(g)	A parent holding company or control person in accordance with § 240.13d-1(b)(1)(ii)(G);
(h)	A savings associations as defined in Section 3(b) of the Federal Deposit Insurance Act (12 U.S.C. 1813);
(i)	A church plan that is excluded from the definition of an investment company under section 3(c)(14) of the Investment Company Act of 1940 (15 U.S.C. 80a-3);
(j)	A non-U.S. institution in accordance with § 240.13d-1(b)(1)(ii)(J). If filing as a non-U.S. institution in accordance with § 240.13d-1(b)(1)(ii)(J), please specify the type of institution:
(k)	Group, in accordance with Rule 240.13d-1(b)(1)(ii)(K).
Item 4.	Ownership
(a)	Amount beneficially owned: 5,592,972
(b)	Percent of class: 5.2%
(c)	Number of shares as to which the person has:
	(i) Sole power to vote or to direct the vote: 5,498,680
	(ii) Shared power to vote or to direct the vote: 94,262
	(iii) Sole power to dispose or to direct the disposition of: 5,498,680
	(iv) Shared power to dispose or to direct the disposition of: 94,262
Item 5.	Ownership of 5 Percent or Less of a Class.
	Not Applicable
Item 6.	Ownership of more than 5 Percent on Behalf of Another Person.
	Not Applicable
Item 7.	Identification and Classification of the Subsidiary Which Acquired the Security Being Reported on by the Parent Holding Company or Control Person.
	Not Applicable
Item 8.	Identification and Classification of Members of the Group.
	Not Applicable
Item 9.	Notice of Dissolution of Group.
	Not Applicable

Item 10.

Certifications:

By signing below I certify that, to the best of my knowledge and belief, the securities referred to above were not acquired and are not held for the purpose of or with the effect of changing or influencing the control of the issuer of the securities and were not acquired and are not held in connection with or as a participant in any transaction having that purpose or effect, other than activities solely in connection with a nomination under § 240.14a-11.

SIGNATURE

After reasonable inquiry and to the best of my knowledge and belief, I certify that the information set forth in this statement is true, complete and correct.

LYTTON LAURENCE W

Signature:	/s/ Laurence W Lytton
Name/Title:	Reporting Person
Date:	08/13/2026