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Eli Lilly and Company reports beneficial ownership of 21,995,880 ordinary shares of ProQR Therapeutics N.V., representing 15.6% of the outstanding class of ordinary shares with a nominal value of 0.04 EUR per share.
Eli Lilly holds sole voting power and sole dispositive power over all 21,995,880 shares and reports no shared voting or dispositive power. The filing is signed by Christopher Anderson, Vice President - Leader of Corporate Securities and Assistant Corporate Secretary.
ProQR Therapeutics N.V. director Bart Filius purchased 150,000 Ordinary Shares on July 17, 2026 at $1.7195 per share in a transaction described as a purchase in the open market or a private transaction. After this trade, he directly holds 300,000 Ordinary Shares of the company.
ProQR Therapeutics N.V. is registering the resale, from time to time, of up to 3,030,303 ordinary shares previously issued to a selling shareholder under an April 7, 2026 Share Purchase Agreement. The company is not selling shares in this offering and will receive no proceeds from any resale.
The shares trade on Nasdaq under “PRQR”; the last reported price was $1.81 per share on July 2, 2026. Ordinary shares outstanding were 105,362,551 as of March 31, 2026 and are expected to be 141,121,476 after including June 2026 equity issuances; this is a baseline figure, not the amount being offered.
ProQR is a clinical-stage biotechnology company developing RNA editing therapies using its Axiomer platform, with lead liver and CNS programs such as AX-0810 for cholestatic diseases and additional collaborations with Lilly and Ginkgo Bioworks. As of March 31, 2026, cash and cash equivalents were €81,088 thousand against total debt of €15,733 thousand.
ProQR Therapeutics N.V. has a shareholder group of Netherlands-based entities and portfolio manager Patrick Johan Hendrik Krol reporting holdings of its ordinary shares on a Schedule 13G/A, based on 141,121,476 ordinary shares outstanding.
Aescap Life Sciences directly holds 4,156,878 shares, or about 2.95% of the class, and Aescap Genetics holds 1,310,271 shares, or 0.93%. Inspirational Visions holds 35,411 shares, or 0.03%. Privium Fund Management B.V., as fund manager of Aescap Life Sciences and Aescap Genetics, may be deemed to beneficially own 5,467,149 shares, or 3.87%.
Through his roles with these entities and a direct holding of 26,302 shares, Krol may be deemed to beneficially own 5,528,862 ProQR shares, or approximately 3.92% of the outstanding ordinary shares. All reporting persons are described as having sole voting and sole dispositive power over their respective positions, with no shared authority reported.
ProQR Therapeutics N.V. Chief Executive Officer Daniel Anton de Boer reported an open-market purchase of 150,000 Ordinary Shares of the company on 9 July 2026 at a price of $1.74 per share. Following this transaction, he directly holds 300,000 Ordinary Shares of ProQR Therapeutics.
ProQR Therapeutics N.V. reported that Chief Executive Officer Daniel Anton de Boer bought Ordinary Shares in the company. On July 7, 2026, he made an open-market purchase of 150,000 Ordinary Shares at $1.59 per share. After this transaction, he directly owns 150,000 Ordinary Shares, indicating this was a new position or a reset position as reflected in this filing.
ProQR Therapeutics N.V. filed a Form F-3 shelf prospectus to register for resale up to 3,030,303 ordinary shares held by a selling shareholder pursuant to a Share Purchase Agreement. The registration is for resale only; ProQR will not receive proceeds and the shares may be sold from time to time after effectiveness.
The prospectus states ordinary shares expected after the offering: up to 141,121,476 (based on 105,362,551 outstanding as of March 31, 2026 plus subsequent issuances through June 30, 2026). Trading price reference: last reported Nasdaq sale $1.81 on July 2, 2026.
ProQR Therapeutics N.V. received an updated ownership filing from Dutch investor Adrianus van Herk and affiliated entities. They report beneficial ownership of 15,384,250 ordinary shares, or about 10.9% of ProQR’s ordinary shares outstanding as of June 26, 2026.
The percentage change is due solely to an increase in ProQR’s total shares outstanding following issuances described in a recent Prospectus Supplement, not from new share purchases or sales by the Van Herk group. The group retains sole or shared voting and dispositive power over the same share amount, and reports no transactions in the past 60 days.
ProQR Therapeutics N.V. director Hinsch Gylvin Lykke received a grant of share options covering 14,595 Ordinary Shares. The options have an exercise price of $1.42 per share, based on the closing market price of the company’s ordinary shares on June 1, 2026.
The options were granted on June 2, 2026, with 25% of the underlying shares vesting and becoming exercisable on June 2, 2027. The remaining 75% vest in 12 substantially equal quarterly installments thereafter, contingent on Lykke’s continued service with ProQR.