STOCK TITAN

[Form 4] Ginkgo Bioworks Holdings, Inc. Insider Trading Activity

Filing Impact
(Low)
Filing Sentiment
(Negative)
Form Type
4
Rhea-AI Filing Summary

Nelnet, Inc. (NNI) – Form 4 filed 07/21/2025

Director and >10% owner Angela L. Muhleisen reported a Code G gift of 41,250 Class A shares on 07/17/2025 at $0.00 per share. Her direct holdings fall to 972,199 shares (down c. 4%).

She continues to report ~2.23 million additional shares held indirectly through her late spouse’s estate (1.23 m) and multiple dynasty trusts/GRATs, bringing total reported beneficial ownership to roughly 3.2 million shares. The filing states that 857,353 shares held by her adult son and 869,025 shares held by her adult daughter are no longer included in her reportable holdings.

No open-market sales, purchases, or derivative transactions were disclosed; the activity appears related to estate or family wealth planning and is unlikely to affect Nelnet’s capital structure or liquidity.

Nelnet, Inc. (NNI) – Modulo 4 presentato il 21/07/2025

La direttrice e azionista con oltre il 10% Angela L. Muhleisen ha segnalato un regalo di tipo G di 41.250 azioni di Classe A il 17/07/2025 al prezzo di $0,00 per azione. Le sue posizioni dirette scendono a 972.199 azioni (in calo di circa il 4%).

Continua a segnalare circa 2,23 milioni di azioni aggiuntive detenute indirettamente tramite l’eredità del defunto coniuge (1,23 milioni) e diversi trust dinastici/GRAT, portando la proprietà beneficiaria totale segnalata a circa 3,2 milioni di azioni. La dichiarazione indica che 857.353 azioni detenute dal figlio adulto e 869.025 azioni detenute dalla figlia adulta non sono più incluse nelle sue partecipazioni segnalabili.

Non sono state dichiarate vendite, acquisti o transazioni su derivati sul mercato aperto; l’attività sembra legata a pianificazione patrimoniale o familiare e probabilmente non influirà sulla struttura del capitale o sulla liquidità di Nelnet.

Nelnet, Inc. (NNI) – Formulario 4 presentado el 21/07/2025

La directora y propietaria de más del 10% Angela L. Muhleisen reportó un regalo Código G de 41,250 acciones Clase A el 17/07/2025 a $0.00 por acción. Sus participaciones directas disminuyen a 972,199 acciones (una caída aproximada del 4%).

Continúa reportando aproximadamente 2.23 millones de acciones adicionales mantenidas indirectamente a través del patrimonio de su difunto cónyuge (1.23 millones) y múltiples fideicomisos dinásticos/GRATs, elevando la propiedad beneficiaria total reportada a cerca de 3.2 millones de acciones. El informe indica que 857,353 acciones en manos de su hijo adulto y 869,025 acciones en manos de su hija adulta ya no se incluyen en sus participaciones reportables.

No se divulgaron ventas, compras ni transacciones con derivados en el mercado abierto; la actividad parece estar relacionada con la planificación patrimonial o familiar y es poco probable que afecte la estructura de capital o liquidez de Nelnet.

Nelnet, Inc. (NNI) – 2025년 7월 21일 제출된 Form 4

이사이자 10% 이상 지분 보유자인 Angela L. Muhleisen가 2025년 7월 17일에 코드 G 증여41,250주 클래스 A 주식을 주당 $0.00에 보고했습니다. 그녀의 직접 보유 주식은 약 4% 감소한 972,199주로 줄었습니다.

고인이 된 배우자의 유산(1.23백만 주)과 여러 다이너스티 트러스트/GRAT를 통해 약 223만 주의 추가 간접 보유 주식을 계속 보고하여 총 보고된 실질 소유 주식은 약 320만 주에 이릅니다. 제출서류에는 성인 아들이 보유한 857,353주와 성인 딸이 보유한 869,025주가 더 이상 보고 대상 보유 주식에 포함되지 않는다고 명시되어 있습니다.

공개 시장에서의 매도, 매수 또는 파생상품 거래는 공개되지 않았으며, 이번 활동은 유산 또는 가족 자산 계획과 관련된 것으로 보이며 Nelnet의 자본 구조나 유동성에 영향을 미칠 가능성은 낮습니다.

Nelnet, Inc. (NNI) – Formulaire 4 déposé le 21/07/2025

La directrice et détentrice de plus de 10% Angela L. Muhleisen a déclaré un don de type Code G de 41 250 actions de Classe A le 17/07/2025 à 0,00 $ par action. Ses participations directes diminuent à 972 199 actions (en baisse d’environ 4%).

Elle continue de déclarer environ 2,23 millions d’actions supplémentaires détenues indirectement via la succession de son défunt conjoint (1,23 million) et plusieurs trusts dynastiques/GRAT, portant la propriété bénéficiaire totale déclarée à environ 3,2 millions d’actions. Le dépôt indique que 857 353 actions détenues par son fils adulte et 869 025 actions détenues par sa fille adulte ne sont plus incluses dans ses participations déclarables.

Aucune vente, achat ou transaction sur dérivés sur le marché ouvert n’a été divulguée ; cette activité semble liée à la planification successorale ou familiale et est peu susceptible d’affecter la structure du capital ou la liquidité de Nelnet.

Nelnet, Inc. (NNI) – Formular 4 eingereicht am 21.07.2025

Die Direktorin und Eigentümerin von mehr als 10% Angela L. Muhleisen meldete am 17.07.2025 eine Schenkung nach Code G von 41.250 Class A Aktien zum Preis von 0,00 $ pro Aktie. Ihr direkter Bestand sinkt auf 972.199 Aktien (rund 4% weniger).

Sie berichtet weiterhin von etwa 2,23 Millionen weiteren indirekt gehaltenen Aktien über den Nachlass ihres verstorbenen Ehepartners (1,23 Mio.) und mehrere Dynastie-Trusts/GRATs, wodurch sich der insgesamt gemeldete wirtschaftliche Eigentum auf etwa 3,2 Millionen Aktien beläuft. Die Meldung gibt an, dass 857.353 Aktien, die ihr erwachsener Sohn hält, und 869.025 Aktien, die ihre erwachsene Tochter hält, nicht mehr in ihren meldungspflichtigen Beständen enthalten sind.

Keine Verkäufe, Käufe oder Derivatgeschäfte auf dem offenen Markt wurden offengelegt; die Transaktion scheint im Zusammenhang mit Nachlass- oder Familienvermögensplanung zu stehen und wird voraussichtlich keine Auswirkungen auf die Kapitalstruktur oder Liquidität von Nelnet haben.

Positive
  • None.
Negative
  • Direct stake reduced by 41,250 shares, though total beneficial ownership remains above 3 million shares.

Insights

TL;DR: Gift of 41k shares; no cash sale; minimal market impact.

The Form 4 shows an internal transfer (Code G) rather than a sale, so there is no price signal or liquidity event. Post-gift, Muhleisen still controls over 3 m shares, preserving her >10% ownership position. Because the transaction is non-monetary and does not change insider sentiment toward the stock, the filing is informational rather than market-moving. Short-term trading implications are negligible.

TL;DR: Estate-planning transfer; ownership concentration unchanged.

The shift of 41,250 shares from direct to other family vehicles slightly lowers direct ownership but leaves overall influence intact. Large indirect holdings through trusts and the estate maintain the Muhleisen family’s controlling interest. Investors should note that the son’s and daughter’s holdings are now excluded, improving disclosure clarity. Governance risk profile remains stable; no red flags surface.

Nelnet, Inc. (NNI) – Modulo 4 presentato il 21/07/2025

La direttrice e azionista con oltre il 10% Angela L. Muhleisen ha segnalato un regalo di tipo G di 41.250 azioni di Classe A il 17/07/2025 al prezzo di $0,00 per azione. Le sue posizioni dirette scendono a 972.199 azioni (in calo di circa il 4%).

Continua a segnalare circa 2,23 milioni di azioni aggiuntive detenute indirettamente tramite l’eredità del defunto coniuge (1,23 milioni) e diversi trust dinastici/GRAT, portando la proprietà beneficiaria totale segnalata a circa 3,2 milioni di azioni. La dichiarazione indica che 857.353 azioni detenute dal figlio adulto e 869.025 azioni detenute dalla figlia adulta non sono più incluse nelle sue partecipazioni segnalabili.

Non sono state dichiarate vendite, acquisti o transazioni su derivati sul mercato aperto; l’attività sembra legata a pianificazione patrimoniale o familiare e probabilmente non influirà sulla struttura del capitale o sulla liquidità di Nelnet.

Nelnet, Inc. (NNI) – Formulario 4 presentado el 21/07/2025

La directora y propietaria de más del 10% Angela L. Muhleisen reportó un regalo Código G de 41,250 acciones Clase A el 17/07/2025 a $0.00 por acción. Sus participaciones directas disminuyen a 972,199 acciones (una caída aproximada del 4%).

Continúa reportando aproximadamente 2.23 millones de acciones adicionales mantenidas indirectamente a través del patrimonio de su difunto cónyuge (1.23 millones) y múltiples fideicomisos dinásticos/GRATs, elevando la propiedad beneficiaria total reportada a cerca de 3.2 millones de acciones. El informe indica que 857,353 acciones en manos de su hijo adulto y 869,025 acciones en manos de su hija adulta ya no se incluyen en sus participaciones reportables.

No se divulgaron ventas, compras ni transacciones con derivados en el mercado abierto; la actividad parece estar relacionada con la planificación patrimonial o familiar y es poco probable que afecte la estructura de capital o liquidez de Nelnet.

Nelnet, Inc. (NNI) – 2025년 7월 21일 제출된 Form 4

이사이자 10% 이상 지분 보유자인 Angela L. Muhleisen가 2025년 7월 17일에 코드 G 증여41,250주 클래스 A 주식을 주당 $0.00에 보고했습니다. 그녀의 직접 보유 주식은 약 4% 감소한 972,199주로 줄었습니다.

고인이 된 배우자의 유산(1.23백만 주)과 여러 다이너스티 트러스트/GRAT를 통해 약 223만 주의 추가 간접 보유 주식을 계속 보고하여 총 보고된 실질 소유 주식은 약 320만 주에 이릅니다. 제출서류에는 성인 아들이 보유한 857,353주와 성인 딸이 보유한 869,025주가 더 이상 보고 대상 보유 주식에 포함되지 않는다고 명시되어 있습니다.

공개 시장에서의 매도, 매수 또는 파생상품 거래는 공개되지 않았으며, 이번 활동은 유산 또는 가족 자산 계획과 관련된 것으로 보이며 Nelnet의 자본 구조나 유동성에 영향을 미칠 가능성은 낮습니다.

Nelnet, Inc. (NNI) – Formulaire 4 déposé le 21/07/2025

La directrice et détentrice de plus de 10% Angela L. Muhleisen a déclaré un don de type Code G de 41 250 actions de Classe A le 17/07/2025 à 0,00 $ par action. Ses participations directes diminuent à 972 199 actions (en baisse d’environ 4%).

Elle continue de déclarer environ 2,23 millions d’actions supplémentaires détenues indirectement via la succession de son défunt conjoint (1,23 million) et plusieurs trusts dynastiques/GRAT, portant la propriété bénéficiaire totale déclarée à environ 3,2 millions d’actions. Le dépôt indique que 857 353 actions détenues par son fils adulte et 869 025 actions détenues par sa fille adulte ne sont plus incluses dans ses participations déclarables.

Aucune vente, achat ou transaction sur dérivés sur le marché ouvert n’a été divulguée ; cette activité semble liée à la planification successorale ou familiale et est peu susceptible d’affecter la structure du capital ou la liquidité de Nelnet.

Nelnet, Inc. (NNI) – Formular 4 eingereicht am 21.07.2025

Die Direktorin und Eigentümerin von mehr als 10% Angela L. Muhleisen meldete am 17.07.2025 eine Schenkung nach Code G von 41.250 Class A Aktien zum Preis von 0,00 $ pro Aktie. Ihr direkter Bestand sinkt auf 972.199 Aktien (rund 4% weniger).

Sie berichtet weiterhin von etwa 2,23 Millionen weiteren indirekt gehaltenen Aktien über den Nachlass ihres verstorbenen Ehepartners (1,23 Mio.) und mehrere Dynastie-Trusts/GRATs, wodurch sich der insgesamt gemeldete wirtschaftliche Eigentum auf etwa 3,2 Millionen Aktien beläuft. Die Meldung gibt an, dass 857.353 Aktien, die ihr erwachsener Sohn hält, und 869.025 Aktien, die ihre erwachsene Tochter hält, nicht mehr in ihren meldungspflichtigen Beständen enthalten sind.

Keine Verkäufe, Käufe oder Derivatgeschäfte auf dem offenen Markt wurden offengelegt; die Transaktion scheint im Zusammenhang mit Nachlass- oder Familienvermögensplanung zu stehen und wird voraussichtlich keine Auswirkungen auf die Kapitalstruktur oder Liquidität von Nelnet haben.

SEC Form 4
FORM 4 UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934
or Section 30(h) of the Investment Company Act of 1940
OMB APPROVAL
OMB Number: 3235-0287
Estimated average burden
hours per response: 0.5
Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).
1. Name and Address of Reporting Person*
Coen Steven P.

(Last) (First) (Middle)
C/O GINKGO BIOWORKS HOLDINGS, INC.
27 DRYDOCK AVENUE

(Street)
BOSTON MA 02210

(City) (State) (Zip)
2. Issuer Name and Ticker or Trading Symbol
Ginkgo Bioworks Holdings, Inc. [ DNA ]
5. Relationship of Reporting Person(s) to Issuer
(Check all applicable)
Director 10% Owner
X Officer (give title below) Other (specify below)
See remarks
3. Date of Earliest Transaction (Month/Day/Year)
07/16/2025
4. If Amendment, Date of Original Filed (Month/Day/Year)
6. Individual or Joint/Group Filing (Check Applicable Line)
X Form filed by One Reporting Person
Form filed by More than One Reporting Person
Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1. Title of Security (Instr. 3) 2. Transaction Date (Month/Day/Year) 2A. Deemed Execution Date, if any (Month/Day/Year) 3. Transaction Code (Instr. 8) 4. Securities Acquired (A) or Disposed Of (D) (Instr. 3, 4 and 5) 5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4) 6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4) 7. Nature of Indirect Beneficial Ownership (Instr. 4)
Code V Amount (A) or (D) Price
Class A Common Stock 07/16/2025 M(1) 1,174 A (1) 10,390 D
Class A Common Stock 07/16/2025 M(1) 312 A (1) 10,702 D
Class A Common Stock 07/16/2025 M(1) 1,061 A (1) 11,763 D
Class A Common Stock 07/17/2025 S(2) 1,084 D $9.842 10,679 D
Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned
(e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security (Instr. 3) 2. Conversion or Exercise Price of Derivative Security 3. Transaction Date (Month/Day/Year) 3A. Deemed Execution Date, if any (Month/Day/Year) 4. Transaction Code (Instr. 8) 5. Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5) 6. Date Exercisable and Expiration Date (Month/Day/Year) 7. Title and Amount of Securities Underlying Derivative Security (Instr. 3 and 4) 8. Price of Derivative Security (Instr. 5) 9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4) 10. Ownership Form: Direct (D) or Indirect (I) (Instr. 4) 11. Nature of Indirect Beneficial Ownership (Instr. 4)
Code V (A) (D) Date Exercisable Expiration Date Title Amount or Number of Shares
Restricted Stock Units (1) 07/16/2025 M(1) 1,174 (3) (3) Class A Common Stock 1,174 (1) 12,927 D
Restricted Stock Units (1) 07/16/2025 M(1) 312 (4) (4) Class A Common Stock 312 (1) 5,003 D
Restricted Stock Units (1) 07/16/2025 M(1) 1,061 (5) (5) Class A Common Stock 1,061 (1) 2,123 D
Explanation of Responses:
1. Each restricted stock unit ("RSU") represents a contingent right to receive one share of the Issuer's Class A Common Stock.
2. Represents shares sold by the Reporting Person to cover tax withholding obligations in connection with the vesting of restricted stock and/or restricted stock units. Sales to cover tax withholding obligations in connection with the vesting of such securities do not represent discretionary trades by the Reporting Person. The Issuer's equity incentive plans allow the Issuer to require that satisfaction of tax withholding obligations be funded by a "sell to cover" transaction.
3. The RSUs vest as follows: 25% of the underlying shares vested on May 1, 2024, then 36 equal monthly installments thereafter.
4. The RSUs vest as follows: 2/48ths of the underlying shares vested on May 1, 2024, then 46 equal monthly installments thereafter.
5. The RSUs vest as follows: 25% vested on April 16, 2025; 25% vested on July 16, 2025; 25% will vest on October 16, 2025; and 25% will vest on January 16, 2026.
Remarks:
Chief Financial Officer and Chief Accounting Officer
/s/ Karen Tepichin, Attorney-in-Fact 07/21/2025
** Signature of Reporting Person Date
Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
* If the form is filed by more than one reporting person, see Instruction 4 (b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
Note: File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB Number.

FAQ

How many Nelnet (NNI) shares did Angela L. Muhleisen transfer?

She gifted 41,250 Class A shares on 07/17/2025.

What is Muhleisen’s direct Nelnet shareholding after the gift?

Her direct ownership stands at 972,199 shares.

Does the filing involve any stock sales or purchases?

No. The only reported transaction is a gift (Code G) with no cash proceeds.

How many Nelnet shares does Muhleisen own in total?

Including indirect interests, she reports about 3.2 million shares.

Why were shares held by her adult children removed from the report?

The filing notes Muhleisen no longer has a reportable beneficial interest in those shares, so they are excluded.
Ginkgo Bioworks Holdings Inc

NYSE:DNA

DNA Rankings

DNA Latest News

DNA Latest SEC Filings

DNA Stock Data

581.81M
54.73M
6.6%
79.85%
15.62%
Biotechnology
Biological Products, (no Disgnostic Substances)
Link
United States
BOSTON