Codexis director De Vre sells 1,088 shares at $1.11

The reported sale covered tax liability incurred due to the vesting and release of restricted stock units.

Available on EDGAR 11/10/2026 - 04:07 PM EST Accepted by SEC EDGAR 11/10/2026 - 04:05 PM EST Learn about SEC filing dates

Rhea-AI Filing Summary

Codexis, Inc. (CDXS) director Raymond De Vre sold 1,088 shares of common stock at $1.11 per share on November 9, 2026. The sale was effected to cover tax liability incurred due to the vesting and release of restricted stock units. His reported holdings after the transaction were 83,723 shares, including 15,948 RSUs; no Rule 10b5-1 plan is reported.

Insider Trade Summary

Net Seller: 1,088 shares

Net Sell 1 txn

Insider De Vre Raymond

Role Director

Sold 1,088 shs ($1K)

Share of Position Sold 1.3%

Type	Security	Shares	Price	Value
Sale	Common Stock F1, F2	1,088	$1.11	$1K

Holdings After Transaction: Common Stock — 83,723 shares (Direct)

Holdings Before and After: Common Stock, 84,811 to 83,723 shares (Direct)

Other Insiders: 2 other insiders reported open-market sales of this stock in the 30 days up to this filing.

Footnotes (2)

1. F1. The sale reported on this Form 4 was effected to cover tax liability incurred due to the vesting and release of Restricted Stock Units ("RSUs") held by the Reporting Person.
2. F2. Includes 15,948 RSUs.

Key Figures

Common shares sold: 1,088 shares Sale price: $1.11 per share Shares held after transaction: 83,723 shares +1 more

4 metrics

Common shares sold 1,088 shares November 9, 2026

Sale price $1.11 per share November 9, 2026

Shares held after transaction 83,723 shares Includes 15,948 RSUs

RSUs included in reported holdings 15,948 RSUs Reported holdings after the transaction

Key Terms

Restricted Stock Units, vesting, tax liability

3 terms

Restricted Stock Units financial

"vesting and release of Restricted Stock Units ("RSUs")"

Restricted stock units are a type of company reward where employees are promised shares of stock, but they only fully own these shares after meeting certain conditions, like staying with the company for a set time. They matter because they can become valuable assets and are often used to motivate employees to help the company succeed.

vesting financial

"tax liability incurred due to the vesting and release"

Vesting is the process by which you earn full ownership of something, like company stock or a retirement benefit, over time. It’s like earning the right to keep a gift piece by piece the longer you stay with a company, making sure employees stay committed before they receive all the benefits.

tax liability financial

"cover tax liability incurred due to the vesting and release"

FAQ

AI-generated questions and answers. How Rhea-AI works. Not financial advice.

How many shares did the CDXS director sell, and at what price?

Raymond De Vre, a director of CDXS, sold 1,088 common shares at $1.11 per share on November 9, 2026. The sale was effected to cover tax liability incurred due to vesting and release of RSUs; no Rule 10b5-1 plan is reported. His reported post-transaction holdings were 83,723 shares, including 15,948 RSUs.

AI-generated analysis. How Rhea-AI works. Not financial advice.

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SEC Form 4

FORM 4

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934
or Section 30(h) of the Investment Company Act of 1940

OMB APPROVAL

OMB Number:			3235-0287
Estimated average burden
hours per response:			0.5

Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).

Check this box to indicate that a transaction was made pursuant to a contract, instruction or written plan for the purchase or sale of equity securities of the issuer that is intended to satisfy the affirmative defense conditions of Rule 10b5-1(c). See Instruction 10.

1. Name and Address of Reporting Person^*

De Vre Raymond

(Last) (First) (Middle)

C/O CODEXIS, INC.

200 PENOBSCOT DRIVE

(Street)

REDWOOD CITY CALIFORNIA 94063

(City) (State) (Zip)

UNITED STATES

(Country)

2. Issuer Name and Ticker or Trading Symbol
CODEXIS, INC. [ CDXS ]

5. Relationship of Reporting Person(s) to Issuer
(Check all applicable)

X	Director		10% Owner
	Officer (give title below)		Other (specify below)

2a. Foreign Trading Symbol

3. Date of Earliest Transaction (Month/Day/Year)
11/09/2026

6. Individual or Joint/Group Filing (Check Applicable Line)

X	Form filed by One Reporting Person
	Form filed by More than One Reporting Person

4. If Amendment, Date of Original Filed (Month/Day/Year)

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1. Title of Security (Instr. 3)	2. Transaction Date (Month/Day/Year)	2A. Deemed Execution Date, if any (Month/Day/Year)	3. Transaction Code (Instr. 8)		4. Securities Acquired (A) or Disposed Of (D) (Instr. 3, 4 and 5)			5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4)	6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	7. Nature of Indirect Beneficial Ownership (Instr. 4)
			Code	V	Amount	(A) or (D)	Price
Common Stock	11/09/2026		S(1)		1,088	D	$1.11	83,723(2)	D

Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned (e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security (Instr. 3)	2. Conversion or Exercise Price of Derivative Security	3. Transaction Date (Month/Day/Year)	3A. Deemed Execution Date, if any (Month/Day/Year)	4. Transaction Code (Instr. 8)		5. Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5)		6. Date Exercisable and Expiration Date (Month/Day/Year)		7. Title and Amount of Securities Underlying Derivative Security (Instr. 3 and 4)		8. Price of Derivative Security (Instr. 5)	9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4)	10. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	11. Nature of Indirect Beneficial Ownership (Instr. 4)
				Code	V	(A)	(D)	Date Exercisable	Expiration Date	Title	Amount or Number of Shares

Explanation of Responses:

1. The sale reported on this Form 4 was effected to cover tax liability incurred due to the vesting and release of Restricted Stock Units ("RSUs") held by the Reporting Person.

2. Includes 15,948 RSUs.

	/s/ Georgia Erbez, as Attorney-in-Fact for Raymond De Vre	11/10/2026
	** Signature of Reporting Person	Date
Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
* If the form is filed by more than one reporting person, see Instruction 4 (b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
Note: File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB Number.
* Form 4: SEC 1474 (03-26)