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CODEXIS, INC. (CDXS) is the issuer named in an amended beneficial-ownership report by Aberdeen Group plc and abrdn Inc. Each reports beneficial ownership of 10,797,702 shares, or 9.86%, with shared voting and dispositive power over those shares and no sole voting or dispositive power. abrdn Holdings Limited is identified as the intermediate holding company for abrdn Inc.; the shares are held on behalf of underlying clients.
Key Figures
Beneficial ownership:10,797,702 sharesPercent of class:9.86%Shared voting power:10,797,702 shares+3 more
6 metrics
Beneficial ownership10,797,702 sharesReported by each of Aberdeen Group plc and abrdn Inc.
Percent of class9.86%Reported by each of Aberdeen Group plc and abrdn Inc.
Shared voting power10,797,702 sharesReported by each of Aberdeen Group plc and abrdn Inc.
Shared dispositive power10,797,702 sharesReported by each of Aberdeen Group plc and abrdn Inc.
Sole voting power0 sharesReported by each of Aberdeen Group plc and abrdn Inc.
Sole dispositive power0 sharesReported by each of Aberdeen Group plc and abrdn Inc.
Key Terms
beneficially owned, shared voting power, shared dispositive power
3 terms
beneficially ownedregulatory
"Amount beneficially owned: 10,797,702"
Beneficially owned describes securities or assets where a person has the economic rights and control—such as the right to receive dividends and to direct voting—even if legal title is held in another name. Think of it like having the keys and using a car that’s registered to someone else: you get the benefits and make decisions. Investors care because beneficial ownership reveals who truly controls value and voting power, affecting corporate decisions and takeover dynamics.
shared voting powerregulatory
"Shared voting power: 10,797,702"
Shared voting power occurs when two or more parties jointly have the right to vote or decide how a block of company shares is cast, like co-owners who must agree before moving a piece of furniture. Investors care because who controls voting rights affects board elections, major corporate decisions and takeover outcomes, and shared control can alter regulatory disclosures and the practical influence any holder has over a company’s direction and value.
shared dispositive powerregulatory
"Shared dispositive power: 10,797,702"
FAQ
AI-generated questions and answers. How Rhea-AI works. Not financial advice.
How many CDXS shares do Aberdeen Group plc and abrdn Inc. report?
Aberdeen Group plc and abrdn Inc. each report beneficial ownership of 10,797,702 CDXS shares, or 9.86% of the class. Each reports shared voting and dispositive power over the position.
SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
SCHEDULE 13G
UNDER THE SECURITIES EXCHANGE ACT OF 1934
(Amendment No. 4)
CODEXIS, INC.
(Name of Issuer)
Common Stock, par value $0.0001 per share
(Title of Class of Securities)
192005106
(CUSIP Number)
09/30/2026
(Date of Event Which Requires Filing of this Statement)
Check the appropriate box to designate the rule pursuant to which this Schedule is filed:
Rule 13d-1(b)
Rule 13d-1(c)
Rule 13d-1(d)
schemaVersion:
SCHEDULE 13G
CUSIP Number(s):
192005106
1
Names of Reporting Persons
Aberdeen Group plc
2
Check the appropriate box if a member of a Group (see instructions)
(a)
(b)
3
Sec Use Only
4
Citizenship or Place of Organization
UNITED KINGDOM
Number of Shares Beneficially Owned by Each Reporting Person With:
5
Sole Voting Power
0.00
6
Shared Voting Power
10,797,702.00
7
Sole Dispositive Power
0.00
8
Shared Dispositive Power
10,797,702.00
9
Aggregate Amount Beneficially Owned by Each Reporting Person
10,797,702.00
10
Check box if the aggregate amount in row (9) excludes certain shares (See Instructions)
11
Percent of class represented by amount in row (9)
9.86 %
12
Type of Reporting Person (See Instructions)
HC
SCHEDULE 13G
CUSIP Number(s):
192005106
1
Names of Reporting Persons
abrdn Inc.
2
Check the appropriate box if a member of a Group (see instructions)
(a)
(b)
3
Sec Use Only
4
Citizenship or Place of Organization
UNITED STATES
Number of Shares Beneficially Owned by Each Reporting Person With:
5
Sole Voting Power
0.00
6
Shared Voting Power
10,797,702.00
7
Sole Dispositive Power
0.00
8
Shared Dispositive Power
10,797,702.00
9
Aggregate Amount Beneficially Owned by Each Reporting Person
10,797,702.00
10
Check box if the aggregate amount in row (9) excludes certain shares (See Instructions)
11
Percent of class represented by amount in row (9)
9.86 %
12
Type of Reporting Person (See Instructions)
CO, IA
SCHEDULE 13G
Item 1.
(a)
Name of issuer:
CODEXIS, INC.
(b)
Address of issuer's principal executive offices:
200 PENOBSCOT DRIVE, REDWOOD CITY, CALIFORNIA
94063
Item 2.
(a)
Name of person filing:
Aberdeen Group plc
abrdn Inc.
(b)
Address or principal business office or, if none, residence:
Aberdeen Group plc:
1 George Street
Edinburgh, United Kingdom
EH2 2LL
abrdn Inc.
1900 Market Street Suite 200.
Philadelphia,
PA 19103
(c)
Citizenship:
Aberdeen Group plc - UNITED KINGDOM
abrdn Inc. - UNITED STATES
(d)
Title of class of securities:
Common Stock, par value $0.0001 per share
(e)
CUSIP No.:
192005106
Item 3.
If this statement is filed pursuant to §§ 240.13d-1(b) or 240.13d-2(b) or (c), check whether the person filing is a:
(a)
Broker or dealer registered under section 15 of the Act (15 U.S.C. 78o);
(b)
Bank as defined in section 3(a)(6) of the Act (15 U.S.C. 78c);
(c)
Insurance company as defined in section 3(a)(19) of the Act (15 U.S.C. 78c);
(d)
Investment company registered under section 8 of the Investment Company Act of 1940 (15 U.S.C. 80a-8);
(e)
An investment adviser in accordance with § 240.13d-1(b)(1)(ii)(E);
(f)
An employee benefit plan or endowment fund in accordance with § 240.13d-1(b)(1)(ii)(F);
(g)
A parent holding company or control person in accordance with § 240.13d-1(b)(1)(ii)(G);
(h)
A savings associations as defined in Section 3(b) of the Federal Deposit Insurance Act (12 U.S.C. 1813);
(i)
A church plan that is excluded from the definition of an investment company under section 3(c)(14) of the Investment Company Act of 1940 (15 U.S.C. 80a-3);
(j)
A non-U.S. institution in accordance with § 240.13d-1(b)(1)(ii)(J). If filing as a non-U.S. institution in accordance with § 240.13d-1(b)(1)(ii)(J),
please specify the type of institution:
(k)
Group, in accordance with Rule 240.13d-1(b)(1)(ii)(K).
Item 4.
Ownership
(a)
Amount beneficially owned:
10,797,702
(b)
Percent of class:
9.86 %
(c)
Number of shares as to which the person has:
(i) Sole power to vote or to direct the vote:
Aberdeen Group plc - 0
abrdn Inc. - 0
(ii) Shared power to vote or to direct the vote:
Aberdeen Group plc - 10,797,702
abrdn Inc. - 10,797,702
(iii) Sole power to dispose or to direct the disposition of:
Aberdeen Group plc - 0
abrdn Inc. - 0
(iv) Shared power to dispose or to direct the disposition of:
Aberdeen Group plc - 10,797,702
abrdn Inc. - 10,797,702
Item 5.
Ownership of 5 Percent or Less of a Class.
Not Applicable
Item 6.
Ownership of more than 5 Percent on Behalf of Another Person.
Not Applicable
Item 7.
Identification and Classification of the Subsidiary Which Acquired the Security Being Reported on by the Parent Holding Company or Control Person.
If a parent holding company has filed this schedule, pursuant to Rule 13d-1(b)(ii)(G), so indicate under Item 3(g) and attach an exhibit stating the identity and the Item 3 classification of the relevant subsidiary. If a parent holding company has filed this schedule pursuant to Rule 13d-1(c) or Rule 13d-1(d), attach an exhibit stating the identification of the relevant subsidiary.
See Exhibit
Item 8.
Identification and Classification of Members of the Group.
Not Applicable
Item 9.
Notice of Dissolution of Group.
Not Applicable
Item 10.
Certifications:
By signing below I certify that, to the best of my knowledge and belief, the securities referred to above were acquired and are held in the ordinary course of business and were not acquired and are not held for the purpose of or with the effect of changing or influencing the control of the issuer of the securities and were not acquired and are not held in connection with or as a participant in any transaction having that purpose or effect, other than activities solely in connection with a nomination under § 240.14a-11.
SIGNATURE
After reasonable inquiry and to the best of my knowledge and belief, I certify that the information set forth in this statement is true, complete and correct.
Aberdeen Group plc
Signature:
Frederic Osei
Name/Title:
Frederic Osei, Major Shareholding Reporting Analyst
Date:
09/30/2026
abrdn Inc.
Signature:
Frederic Osei
Name/Title:
Frederic Osei, Major Shareholding Reporting Analyst
Date:
09/30/2026
Exhibit Information
Aberdeen Group plc
abrdn Holdings Limited
abrdn Inc.
Aberdeen Group plc is the parent company.
abrdn Holdings Limited is the intermediate holding company for abrdn Inc.
abrdn Inc. beneficially owns on behalf of our underlying clients 5% or greater of the outstanding shares of the security class being reported on this Schedule 13G.
