Aberdeen Group plc (CDXS) discloses 6.77M-share Codexis stake held via abrdn Inc.

Rhea-AI Filing Summary

Aberdeen Group plc and its subsidiary abrdn Inc. report beneficial ownership of common stock of Codexis, Inc. (par value $0.0001 per share).

They collectively report beneficial ownership of 6,769,954 shares, representing 7.45% of this class. Both entities report no sole voting or dispositive power, but shared voting and shared dispositive power over all 6,769,954 shares. Aberdeen Group plc is the parent of abrdn Holdings Limited, which is the intermediate holding company for abrdn Inc., and abrdn Inc. holds these shares on behalf of underlying clients.

Key Figures

Beneficial ownership: 6,769,954 shares Percent of class: 7.45% Shared voting power: 6,769,954 shares +1 more

4 metrics

Beneficial ownership 6,769,954 shares Common stock of Codexis, Inc. beneficially owned by Aberdeen Group plc and abrdn Inc.

Percent of class 7.45% Percentage of Codexis common stock class beneficially owned

Shared voting power 6,769,954 shares Shares over which Aberdeen Group plc and abrdn Inc. share voting power

Shared dispositive power 6,769,954 shares Shares over which Aberdeen Group plc and abrdn Inc. share dispositive power

Key Terms

beneficially owns, shared voting power, shared dispositive power, parent holding company, +1 more

5 terms

beneficially owns financial

"abrdn Inc. beneficially owns on behalf of our underlying clients 5% or greater"

Beneficially owns means a person or entity enjoys the economic benefits and control of a security even if the legal title or registration is held in another name. Think of it like having the keys and profits from a car that is registered to a friend: you use it, benefit from it, and make decisions about it even though the official paperwork lists someone else. For investors, this matters because it reveals who truly controls shares, affects voting power, potential conflicts of interest, and regulatory disclosure obligations.

shared voting power financial

"Shared Voting Power 6,769,954.00"

Shared voting power occurs when two or more parties jointly have the right to vote or decide how a block of company shares is cast, like co-owners who must agree before moving a piece of furniture. Investors care because who controls voting rights affects board elections, major corporate decisions and takeover outcomes, and shared control can alter regulatory disclosures and the practical influence any holder has over a company’s direction and value.

shared dispositive power financial

"Shared Dispositive Power 6,769,954.00"

parent holding company financial

"If a parent holding company has filed this schedule, pursuant to (ii)(G)"

percent of class financial

"Percent of class: 7.45 %"

Percent of class is the portion of a specific category of securities—such as a company’s common shares, preferred shares, or a bond series—that takes part in or approves a corporate action (vote, consent, tender, etc.). Investors watch this number because it reveals how much support or opposition exists within that particular shareholder group; like counting how many members of a club back a proposal, it can determine whether a plan passes or how influence is distributed.

AI-generated analysis. How Rhea-AI works. Not financial advice.

See more from StockTitan in Google Search and AI answers. Adds StockTitan as a preferred source · opens Google

Add on Google Not now

Available on EDGAR 07/20/2026 - 07:09 AM Accepted by SEC EDGAR 07/20/2026 - 07:08 AM Learn about SEC filing dates

FAQ

How many Codexis (CDXS) shares are beneficially owned by Aberdeen Group and abrdn Inc.?

Aberdeen Group plc and abrdn Inc. report beneficial ownership of 6,769,954 Codexis common shares. This stake represents 7.45% of Codexis’s outstanding common stock of this class, held on behalf of underlying clients.

What percentage of Codexis (CDXS) does abrdn Inc. beneficially own?

abrdn Inc. reports beneficial ownership of 7.45% of Codexis’s common stock class. This corresponds to 6,769,954 shares over which it has shared voting and shared dispositive power, with no sole authority reported.

Do Aberdeen Group and abrdn Inc. have sole or shared voting power over Codexis (CDXS) shares?

They report no sole voting power and shared voting power over 6,769,954 shares of Codexis common stock. They likewise have no sole dispositive power but shared dispositive power over the same number of shares.

Who is the parent company for the Codexis (CDXS) holdings reported by abrdn Inc.?

Aberdeen Group plc is identified as the parent company. abrdn Holdings Limited is the intermediate holding company for abrdn Inc., which beneficially owns the Codexis shares on behalf of underlying clients.

Where are Aberdeen Group plc and abrdn Inc., which hold Codexis (CDXS) shares, organized?

Aberdeen Group plc is organized in the United Kingdom, while abrdn Inc. is organized in the United States. Both entities together report beneficial ownership and shared voting and dispositive power over Codexis shares.

What class of Codexis (CDXS) securities is held by Aberdeen Group and abrdn Inc.?

They hold Codexis common stock with a par value of $0.0001 per share. The reported position is 6,769,954 common shares, representing 7.45% of that security class.

SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

SCHEDULE 13G

UNDER THE SECURITIES EXCHANGE ACT OF 1934

(Amendment No. 3)

CODEXIS, INC.

(Name of Issuer)

Common Stock, par value $0.0001 per share

(Title of Class of Securities)

192005106

(CUSIP Number)

06/30/2026

(Date of Event Which Requires Filing of this Statement)

Check the appropriate box to designate the rule pursuant to which this Schedule is filed:

Rule 13d-1(b)

Rule 13d-1(c)

Rule 13d-1(d)

schemaVersion:

SCHEDULE 13G

CUSIP Number(s): 192005106

1

Names of Reporting Persons

Aberdeen Group plc

2

Check the appropriate box if a member of a Group (see instructions)

(a)
(b)

3

Sec Use Only

4

Citizenship or Place of Organization

UNITED KINGDOM

Number of Shares Beneficially Owned by Each Reporting Person With:

5	Sole Voting Power 0.00
6	Shared Voting Power 6,769,954.00
7	Sole Dispositive Power 0.00
8	Shared Dispositive Power 6,769,954.00

9

Aggregate Amount Beneficially Owned by Each Reporting Person

6,769,954.00

10

Check box if the aggregate amount in row (9) excludes certain shares (See Instructions)

11

Percent of class represented by amount in row (9)

7.45 %

12

Type of Reporting Person (See Instructions)

HC

SCHEDULE 13G

CUSIP Number(s): 192005106

1

Names of Reporting Persons

abrdn Inc.

2

Check the appropriate box if a member of a Group (see instructions)

(a)
(b)

3

Sec Use Only

4

Citizenship or Place of Organization

UNITED STATES

Number of Shares Beneficially Owned by Each Reporting Person With:

5	Sole Voting Power 0.00
6	Shared Voting Power 6,769,954.00
7	Sole Dispositive Power 0.00
8	Shared Dispositive Power 6,769,954.00

9

Aggregate Amount Beneficially Owned by Each Reporting Person

0.00

10

Check box if the aggregate amount in row (9) excludes certain shares (See Instructions)

11

Percent of class represented by amount in row (9)

7.45 %

12

Type of Reporting Person (See Instructions)

CO, IA

SCHEDULE 13G

Item 1.
(a)	Name of issuer: CODEXIS, INC.
(b)	Address of issuer's principal executive offices: 200 PENOBSCOT DRIVE, REDWOOD CITY, CALIFORNIA 94063
Item 2.
(a)	Name of person filing: Aberdeen Group plc abrdn Inc.
(b)	Address or principal business office or, if none, residence: Aberdeen Group plc: 1 George Street Edinburgh, United Kingdom EH2 2LL abrdn Inc. 1900 Market Street Suite 200. Philadelphia, PA 19103
(c)	Citizenship: Aberdeen Group plc - UNITED KINGDOM abrdn Inc. - UNITED STATES
(d)	Title of class of securities: Common Stock, par value $0.0001 per share
(e)	CUSIP No.: 192005106
Item 3.	If this statement is filed pursuant to §§ 240.13d-1(b) or 240.13d-2(b) or (c), check whether the person filing is a:
(a)	Broker or dealer registered under section 15 of the Act (15 U.S.C. 78o);
(b)	Bank as defined in section 3(a)(6) of the Act (15 U.S.C. 78c);
(c)	Insurance company as defined in section 3(a)(19) of the Act (15 U.S.C. 78c);
(d)	Investment company registered under section 8 of the Investment Company Act of 1940 (15 U.S.C. 80a-8);
(e)	An investment adviser in accordance with § 240.13d-1(b)(1)(ii)(E);
(f)	An employee benefit plan or endowment fund in accordance with § 240.13d-1(b)(1)(ii)(F);
(g)	A parent holding company or control person in accordance with § 240.13d-1(b)(1)(ii)(G);
(h)	A savings associations as defined in Section 3(b) of the Federal Deposit Insurance Act (12 U.S.C. 1813);
(i)	A church plan that is excluded from the definition of an investment company under section 3(c)(14) of the Investment Company Act of 1940 (15 U.S.C. 80a-3);
(j)	A non-U.S. institution in accordance with § 240.13d-1(b)(1)(ii)(J). If filing as a non-U.S. institution in accordance with § 240.13d-1(b)(1)(ii)(J), please specify the type of institution:
(k)	Group, in accordance with Rule 240.13d-1(b)(1)(ii)(K).
Item 4.	Ownership
(a)	Amount beneficially owned: 6,769,954
(b)	Percent of class: 7.45 %
(c)	Number of shares as to which the person has:
	(i) Sole power to vote or to direct the vote: Aberdeen Group plc - 0 abrdn Inc. - 0
	(ii) Shared power to vote or to direct the vote: Aberdeen Group plc - 6,769,954 abrdn Inc. - 6,769,954
	(iii) Sole power to dispose or to direct the disposition of: Aberdeen Group plc - 0 abrdn Inc. - 0
	(iv) Shared power to dispose or to direct the disposition of: Aberdeen Group plc - 6,769,954 abrdn Inc. - 6,769,954
Item 5.	Ownership of 5 Percent or Less of a Class.
	Not Applicable
Item 6.	Ownership of more than 5 Percent on Behalf of Another Person.
	Not Applicable
Item 7.	Identification and Classification of the Subsidiary Which Acquired the Security Being Reported on by the Parent Holding Company or Control Person.
	If a parent holding company has filed this schedule, pursuant to Rule 13d-1(b)(ii)(G), so indicate under Item 3(g) and attach an exhibit stating the identity and the Item 3 classification of the relevant subsidiary. If a parent holding company has filed this schedule pursuant to Rule 13d-1(c) or Rule 13d-1(d), attach an exhibit stating the identification of the relevant subsidiary. See Exhibit
Item 8.	Identification and Classification of Members of the Group.
	Not Applicable
Item 9.	Notice of Dissolution of Group.
	Not Applicable

Item 10.

Certifications:

By signing below I certify that, to the best of my knowledge and belief, the securities referred to above were acquired and are held in the ordinary course of business and were not acquired and are not held for the purpose of or with the effect of changing or influencing the control of the issuer of the securities and were not acquired and are not held in connection with or as a participant in any transaction having that purpose or effect, other than activities solely in connection with a nomination under § 240.14a-11.

SIGNATURE

After reasonable inquiry and to the best of my knowledge and belief, I certify that the information set forth in this statement is true, complete and correct.

Aberdeen Group plc

Signature:	Lyn Beaton
Name/Title:	Lyn Beaton, Senior Major Shareholding Reporting Analyst
Date:	07/15/2026

abrdn Inc.

Signature:	Lyn Beaton
Name/Title:	Lyn Beaton, Senior Major Shareholding Reporting Analyst
Date:	07/15/2026

Exhibit Information

Aberdeen Group plc abrdn Holdings Limited abrdn Inc. Aberdeen Group plc is the parent company. abrdn Holdings Limited is the intermediate holding company for abrdn Inc. abrdn Inc. beneficially owns on behalf of our underlying clients 5% or greater of the outstanding shares of the security class being reported on this Schedule 13G.