abrdn reports 8.82% Codexis (CDXS) stake in Schedule 13G/A filing update

Rhea-AI Filing Summary

Aberdeen Group plc and abrdn Inc. filed an amended Schedule 13G reporting their beneficial ownership of 7,966,498 shares of Codexis, Inc. common stock, representing 8.82% of the class as of 12/31/2025. All of these shares are reported with shared voting and shared dispositive power, with no sole voting or dispositive authority.

The filing states that the securities were acquired and are held in the ordinary course of business and not for the purpose of changing or influencing control of Codexis. abrdn Inc. holds the position on behalf of underlying clients, with Aberdeen Group plc as the parent company and abrdn Holdings Limited as an intermediate holding company.

SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

SCHEDULE 13G

UNDER THE SECURITIES EXCHANGE ACT OF 1934

(Amendment No. 2)

CODEXIS, INC.

(Name of Issuer)

Common Stock, par value $0.0001 per share

(Title of Class of Securities)

192005106

(CUSIP Number)

12/31/2025

(Date of Event Which Requires Filing of this Statement)

Check the appropriate box to designate the rule pursuant to which this Schedule is filed:

Rule 13d-1(b)

Rule 13d-1(c)

Rule 13d-1(d)

SCHEDULE 13G

CUSIP No. 192005106

1

Names of Reporting Persons

Aberdeen Group plc

2

Check the appropriate box if a member of a Group (see instructions)

(a)
(b)

3

Sec Use Only

4

Citizenship or Place of Organization

UNITED KINGDOM

Number of Shares Beneficially Owned by Each Reporting Person With:

5	Sole Voting Power 0.00
6	Shared Voting Power 7,966,498.00
7	Sole Dispositive Power 0.00
8	Shared Dispositive Power 7,966,498.00

9

Aggregate Amount Beneficially Owned by Each Reporting Person

7,966,498.00

10

Check box if the aggregate amount in row (9) excludes certain shares (See Instructions)

11

Percent of class represented by amount in row (9)

8.82 %

12

Type of Reporting Person (See Instructions)

HC

SCHEDULE 13G

CUSIP No. 192005106

1

Names of Reporting Persons

abrdn Inc.

2

Check the appropriate box if a member of a Group (see instructions)

(a)
(b)

3

Sec Use Only

4

Citizenship or Place of Organization

UNITED STATES

Number of Shares Beneficially Owned by Each Reporting Person With:

5	Sole Voting Power 0.00
6	Shared Voting Power 7,966,498.00
7	Sole Dispositive Power 0.00
8	Shared Dispositive Power 7,966,498.00

9

Aggregate Amount Beneficially Owned by Each Reporting Person

7,966,498.00

10

Check box if the aggregate amount in row (9) excludes certain shares (See Instructions)

11

Percent of class represented by amount in row (9)

8.82 %

12

Type of Reporting Person (See Instructions)

CO, IA

SCHEDULE 13G

Item 1.
(a)	Name of issuer: CODEXIS, INC.
(b)	Address of issuer's principal executive offices: 200 PENOBSCOT DRIVE, REDWOOD CITY, CALIFORNIA 94063
Item 2.
(a)	Name of person filing: Aberdeen Group plc abrdn Inc.
(b)	Address or principal business office or, if none, residence: Aberdeen Group plc: 1 George Street Edinburgh, United Kingdom EH2 2LL abrdn Inc. 1900 Market Street Suite 200. Philadelphia, PA 19103
(c)	Citizenship: Aberdeen Group plc - UNITED KINGDOM abrdn Inc. - UNITED STATES
(d)	Title of class of securities: Common Stock, par value $0.0001 per share
(e)	CUSIP No.: 192005106
Item 3.	If this statement is filed pursuant to §§ 240.13d-1(b) or 240.13d-2(b) or (c), check whether the person filing is a:
(a)	Broker or dealer registered under section 15 of the Act (15 U.S.C. 78o);
(b)	Bank as defined in section 3(a)(6) of the Act (15 U.S.C. 78c);
(c)	Insurance company as defined in section 3(a)(19) of the Act (15 U.S.C. 78c);
(d)	Investment company registered under section 8 of the Investment Company Act of 1940 (15 U.S.C. 80a-8);
(e)	An investment adviser in accordance with § 240.13d-1(b)(1)(ii)(E);
(f)	An employee benefit plan or endowment fund in accordance with § 240.13d-1(b)(1)(ii)(F);
(g)	A parent holding company or control person in accordance with § 240.13d-1(b)(1)(ii)(G);
(h)	A savings associations as defined in Section 3(b) of the Federal Deposit Insurance Act (12 U.S.C. 1813);
(i)	A church plan that is excluded from the definition of an investment company under section 3(c)(14) of the Investment Company Act of 1940 (15 U.S.C. 80a-3);
(j)	A non-U.S. institution in accordance with § 240.13d-1(b)(1)(ii)(J). If filing as a non-U.S. institution in accordance with § 240.13d-1(b)(1)(ii)(J), please specify the type of institution:
(k)	Group, in accordance with Rule 240.13d-1(b)(1)(ii)(K).
Item 4.	Ownership
(a)	Amount beneficially owned: 7,966,498
(b)	Percent of class: 8.82 %
(c)	Number of shares as to which the person has:
	(i) Sole power to vote or to direct the vote: Aberdeen Group plc - 0 abrdn Inc. - 0
	(ii) Shared power to vote or to direct the vote: Aberdeen Group plc - 7,966,498 abrdn Inc. - 7,966,498
	(iii) Sole power to dispose or to direct the disposition of: Aberdeen Group plc - 0 abrdn Inc. - 0
	(iv) Shared power to dispose or to direct the disposition of: Aberdeen Group plc - 7,966,498 abrdn Inc. - 7,966,498
Item 5.	Ownership of 5 Percent or Less of a Class.
	Not Applicable
Item 6.	Ownership of more than 5 Percent on Behalf of Another Person.
	Not Applicable
Item 7.	Identification and Classification of the Subsidiary Which Acquired the Security Being Reported on by the Parent Holding Company or Control Person.
	If a parent holding company has filed this schedule, pursuant to Rule 13d-1(b)(ii)(G), so indicate under Item 3(g) and attach an exhibit stating the identity and the Item 3 classification of the relevant subsidiary. If a parent holding company has filed this schedule pursuant to Rule 13d-1(c) or Rule 13d-1(d), attach an exhibit stating the identification of the relevant subsidiary. See Exhibit
Item 8.	Identification and Classification of Members of the Group.
	Not Applicable
Item 9.	Notice of Dissolution of Group.
	Not Applicable

Item 10.

Certifications:

By signing below I certify that, to the best of my knowledge and belief, the securities referred to above were acquired and are held in the ordinary course of business and were not acquired and are not held for the purpose of or with the effect of changing or influencing the control of the issuer of the securities and were not acquired and are not held in connection with or as a participant in any transaction having that purpose or effect, other than activities solely in connection with a nomination under ?? 240.14a-11.

SIGNATURE

After reasonable inquiry and to the best of my knowledge and belief, I certify that the information set forth in this statement is true, complete and correct.

Aberdeen Group plc

Signature:	Cara Bird
Name/Title:	Cara Bird, Major Shareholding Reporting Analyst
Date:	01/15/2026

abrdn Inc.

Signature:	Cara Bird
Name/Title:	Cara Bird, Major Shareholding Reporting Analyst
Date:	01/15/2026

Exhibit Information

Aberdeen Group plc abrdn Holdings Limited abrdn Inc. Aberdeen Group plc is the parent company. abrdn Holdings Limited is the intermediate holding company for abrdn Inc. abrdn Inc. beneficially owns on behalf of our underlying clients 5% or greater of the outstanding shares of the security class being reported on this Schedule 13G.

FAQ

What percentage of Codexis (CDXS) does abrdn report owning?

Aberdeen Group plc and abrdn Inc. report beneficial ownership of 8.82% of Codexis, Inc.'s common stock, based on 7,966,498 shares.

How many Codexis shares are beneficially owned according to this Schedule 13G/A?

The reporting persons state they beneficially own 7,966,498 shares of Codexis, Inc. common stock.

Who are the reporting persons in this Codexis Schedule 13G/A filing?

The reporting persons are Aberdeen Group plc and abrdn Inc., with Aberdeen Group plc as the parent company and abrdn Holdings Limited as the intermediate holding company for abrdn Inc.

Do Aberdeen Group plc and abrdn Inc. seek to influence control of Codexis?

No. They certify that the securities were acquired and are held in the ordinary course of business and not for the purpose of changing or influencing the control of Codexis, Inc.

What voting and dispositive powers do the reporting persons have over Codexis shares?

Both Aberdeen Group plc and abrdn Inc. report 0 shares with sole voting or dispositive power and 7,966,498 shares with shared voting and shared dispositive power.

As of what date is the Codexis ownership information reported in this Schedule 13G/A?

The ownership information is reported as of 12/31/2025, which is the date of the event requiring the filing.

On whose behalf does abrdn Inc. hold the Codexis shares?

According to the exhibit, abrdn Inc. beneficially owns the Codexis shares on behalf of its underlying clients who hold 5% or more of the outstanding shares of the reported security class.