Laurence Lytton holds 4.53M shares at Codexis (NASDAQ: CDXS)

Rhea-AI Filing Summary

Codexis, Inc. beneficial owner Laurence W. Lytton reports beneficial ownership of 4,532,069 shares of Common Stock, representing 5.0%.

The percentage is calculated based on 90,324,383 shares outstanding as of November 3, 2025, per the issuer's reported Form 10-Q. The filing is a Schedule 13G disclosure of ownership.

Insights

Corporate Governance Analyst

Beneficial ownership disclosure shows a passive 5.0% stake by Laurence Lytton.

The filing lists 4,532,069 shares beneficially owned with sole voting and dispositive power. This Schedule 13G format typically indicates passive or passive-investor status rather than an active trading disclosure.

Ownership is sized at 5.0% of the reported 90,324,383 share base as of November 3, 2025; subsequent filings may update this position.

SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

SCHEDULE 13G

UNDER THE SECURITIES EXCHANGE ACT OF 1934

Codexis, Inc.

(Name of Issuer)

Common Stock

(Title of Class of Securities)

03/06/2026

(Date of Event Which Requires Filing of this Statement)

Check the appropriate box to designate the rule pursuant to which this Schedule is filed:

Rule 13d-1(b)

Rule 13d-1(c)

Rule 13d-1(d)

SCHEDULE 13G

CUSIP Number(s):

1

Names of Reporting Persons

LYTTON LAURENCE W

2

Check the appropriate box if a member of a Group (see instructions)

(a)
(b)

3

Sec Use Only

4

Citizenship or Place of Organization

UNITED STATES

Number of Shares Beneficially Owned by Each Reporting Person With:

5	Sole Voting Power 4,532,069.00
6	Shared Voting Power 0.00
7	Sole Dispositive Power 4,532,069.00
8	Shared Dispositive Power 0.00

9

Aggregate Amount Beneficially Owned by Each Reporting Person

4,532,069.00

10

Check box if the aggregate amount in row (9) excludes certain shares (See Instructions)

11

Percent of class represented by amount in row (9)

5.0 %

12

Type of Reporting Person (See Instructions)

IN

Comment for Type of Reporting Person: Percentage calculated based on 90,324,383 shares of Common Stock outstanding on November 3, 2025, as reported in the Form 10-Q filed by the Issuer for the quarterly period ended September 30, 2025.

SCHEDULE 13G

Item 1.
(a)	Name of issuer: Codexis, Inc.
(b)	Address of issuer's principal executive offices: 200 Penobscot Drive, Redwood City, CA 94063
Item 2.
(a)	Name of person filing: Laurence W. Lytton
(b)	Address or principal business office or, if none, residence: 467 Central Park West New York, NY 10025
(c)	Citizenship: USA
(d)	Title of class of securities: Common Stock
(e)	CUSIP No.: 192005106
Item 3.	If this statement is filed pursuant to §§ 240.13d-1(b) or 240.13d-2(b) or (c), check whether the person filing is a:
(a)	Broker or dealer registered under section 15 of the Act (15 U.S.C. 78o);
(b)	Bank as defined in section 3(a)(6) of the Act (15 U.S.C. 78c);
(c)	Insurance company as defined in section 3(a)(19) of the Act (15 U.S.C. 78c);
(d)	Investment company registered under section 8 of the Investment Company Act of 1940 (15 U.S.C. 80a-8);
(e)	An investment adviser in accordance with § 240.13d-1(b)(1)(ii)(E);
(f)	An employee benefit plan or endowment fund in accordance with § 240.13d-1(b)(1)(ii)(F);
(g)	A parent holding company or control person in accordance with § 240.13d-1(b)(1)(ii)(G);
(h)	A savings associations as defined in Section 3(b) of the Federal Deposit Insurance Act (12 U.S.C. 1813);
(i)	A church plan that is excluded from the definition of an investment company under section 3(c)(14) of the Investment Company Act of 1940 (15 U.S.C. 80a-3);
(j)	A non-U.S. institution in accordance with § 240.13d-1(b)(1)(ii)(J). If filing as a non-U.S. institution in accordance with § 240.13d-1(b)(1)(ii)(J), please specify the type of institution:
(k)	Group, in accordance with Rule 240.13d-1(b)(1)(ii)(K).
Item 4.	Ownership
(a)	Amount beneficially owned: 4,532,069
(b)	Percent of class: 5.0%
(c)	Number of shares as to which the person has:
	(i) Sole power to vote or to direct the vote: 4,532,069
	(ii) Shared power to vote or to direct the vote: 0
	(iii) Sole power to dispose or to direct the disposition of: 4,532,069
	(iv) Shared power to dispose or to direct the disposition of: 0
Item 5.	Ownership of 5 Percent or Less of a Class.
	Not Applicable
Item 6.	Ownership of more than 5 Percent on Behalf of Another Person.
	Not Applicable
Item 7.	Identification and Classification of the Subsidiary Which Acquired the Security Being Reported on by the Parent Holding Company or Control Person.
	Not Applicable
Item 8.	Identification and Classification of Members of the Group.
	Not Applicable
Item 9.	Notice of Dissolution of Group.
	Not Applicable

Item 10.

Certifications:

By signing below I certify that, to the best of my knowledge and belief, the securities referred to above were not acquired and are not held for the purpose of or with the effect of changing or influencing the control of the issuer of the securities and were not acquired and are not held in connection with or as a participant in any transaction having that purpose or effect, other than activities solely in connection with a nomination under ?? 240.14a-11.

SIGNATURE

After reasonable inquiry and to the best of my knowledge and belief, I certify that the information set forth in this statement is true, complete and correct.

LYTTON LAURENCE W

Signature:	/s/ Laurence W Lytton
Name/Title:	Reporting Person
Date:	03/11/2026

FAQ

What stake does Laurence W. Lytton report in Codexis (CDXS)?

He reports beneficial ownership of 4,532,069 shares, representing 5.0%. The percentage uses a share base of 90,324,383 outstanding as of November 3, 2025, per the company's Form 10-Q.

Does the Schedule 13G indicate voting or dispositive power for Lytton?

Yes. The filing shows sole voting power and sole dispositive power over 4,532,069 shares. There is no shared voting or dispositive power reported in the filing.

How was the 5.0% ownership percentage calculated?

The percentage is calculated using 90,324,383 shares outstanding as reported in the issuer's Form 10-Q for the quarter ended September 30, 2025. That base yields the reported 5.0% figure.

Is this Schedule 13G an indication of active trading by the holder?

A Schedule 13G typically reflects a passive beneficial ownership disclosure rather than active trading. The filing format and statements here align with passive reporting rather than Form 13D active-acquirer disclosure.

When was the Schedule 13G signed and filed by the reporting person?

The filing is signed by Laurence W. Lytton with a signature date of 03/11/2026. The ownership data references the outstanding share count as of November 3, 2025 in the supporting note.