Aberdeen Group files 13G for 4.2M Codexis shares, hitting 5% threshold

Rhea-AI Filing Summary

Aberdeen Group plc and its U.S. affiliate abrdn Inc. jointly filed a Schedule 13G reporting passive ownership of 4,200,950 Codexis (CDXS) common shares as of 30 Jun 2025. The position equals 5.07 % of outstanding stock, making the firms a 5-percent beneficial owner under SEC rules.

The filing states shared voting and dispositive power over the entire stake; neither entity holds sole authority. Aberdeen is classified as a parent holding company/control person (HC) incorporated in the United Kingdom, while abrdn Inc. is a U.S. corporation and registered investment adviser (CO, IA). Both certify the shares were acquired in the ordinary course of business and not to influence control of Codexis.

Signatures were provided by Major Shareholding Reporting Analyst Jenny Clarke on 30 Jul 2025. No exhibits indicate activist intent, group formation, or additional derivatives. The disclosure primarily informs investors of a new institutional holder exceeding the 5 % threshold, adding visibility to Codexis’ shareholder base without signaling governance changes.

Positive

• abrdn/Aberdeen disclose a 5.07 % passive stake, adding a well-known institutional investor to Codexis’ shareholder base and potentially supporting stock liquidity.

Negative

• None.

Insights

Securities Analyst

TL;DR: abrdn now owns 5.07 % of CDXS; passive stake raises institutional profile but offers limited strategic influence.

This Schedule 13G is informational rather than catalytic. abrdn’s 4.2 M-share position makes it a top holder for a sub-$1 bn market-cap company, potentially improving liquidity and signaling confidence in Codexis’ prospects. However, the lack of sole voting power and the Rule 13d-1(b) passive filing status suggest no immediate push for operational or governance changes. The impact on valuation depends on whether other institutions follow; as it stands, it modestly broadens institutional ownership without affecting control dynamics or capital structure.

SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

SCHEDULE 13G

UNDER THE SECURITIES EXCHANGE ACT OF 1934

CODEXIS, INC.

(Name of Issuer)

Common Stock, par value $0.0001 per share

(Title of Class of Securities)

192005106

(CUSIP Number)

06/30/2025

(Date of Event Which Requires Filing of this Statement)

Check the appropriate box to designate the rule pursuant to which this Schedule is filed:

Rule 13d-1(b)

Rule 13d-1(c)

Rule 13d-1(d)

SCHEDULE 13G

CUSIP No. 192005106

1

Names of Reporting Persons

Aberdeen Group plc

2

Check the appropriate box if a member of a Group (see instructions)

(a)
(b)

3

Sec Use Only

4

Citizenship or Place of Organization

UNITED KINGDOM

Number of Shares Beneficially Owned by Each Reporting Person With:

5	Sole Voting Power 0.00
6	Shared Voting Power 4,200,950.00
7	Sole Dispositive Power 0.00
8	Shared Dispositive Power 4,200,950.00

9

Aggregate Amount Beneficially Owned by Each Reporting Person

4,200,950.00

10

Check box if the aggregate amount in row (9) excludes certain shares (See Instructions)

11

Percent of class represented by amount in row (9)

5.07 %

12

Type of Reporting Person (See Instructions)

HC

SCHEDULE 13G

CUSIP No. 192005106

1

Names of Reporting Persons

abrdn Inc.

2

Check the appropriate box if a member of a Group (see instructions)

(a)
(b)

3

Sec Use Only

4

Citizenship or Place of Organization

UNITED STATES

Number of Shares Beneficially Owned by Each Reporting Person With:

5	Sole Voting Power 0.00
6	Shared Voting Power 4,200,950.00
7	Sole Dispositive Power 0.00
8	Shared Dispositive Power 4,200,950.00

9

Aggregate Amount Beneficially Owned by Each Reporting Person

4,200,950.00

10

Check box if the aggregate amount in row (9) excludes certain shares (See Instructions)

11

Percent of class represented by amount in row (9)

5.07 %

12

Type of Reporting Person (See Instructions)

CO, IA

SCHEDULE 13G

Item 1.
(a)	Name of issuer: CODEXIS, INC.
(b)	Address of issuer's principal executive offices: 200 PENOBSCOT DRIVE, REDWOOD CITY, CALIFORNIA 94063
Item 2.
(a)	Name of person filing: Aberdeen Group plc abrdn Inc.
(b)	Address or principal business office or, if none, residence: Aberdeen Group plc: 1 George Street Edinburgh, United Kingdom EH2 2LL abrdn Inc. 1900 Market Street Suite 200. Philadelphia, PA 19103
(c)	Citizenship: Aberdeen Group plc - UNITED KINGDOM abrdn Inc. - UNITED STATES
(d)	Title of class of securities: Common Stock, par value $0.0001 per share
(e)	CUSIP No.: 192005106
Item 3.	If this statement is filed pursuant to §§ 240.13d-1(b) or 240.13d-2(b) or (c), check whether the person filing is a:
(a)	Broker or dealer registered under section 15 of the Act (15 U.S.C. 78o);
(b)	Bank as defined in section 3(a)(6) of the Act (15 U.S.C. 78c);
(c)	Insurance company as defined in section 3(a)(19) of the Act (15 U.S.C. 78c);
(d)	Investment company registered under section 8 of the Investment Company Act of 1940 (15 U.S.C. 80a-8);
(e)	An investment adviser in accordance with § 240.13d-1(b)(1)(ii)(E);
(f)	An employee benefit plan or endowment fund in accordance with § 240.13d-1(b)(1)(ii)(F);
(g)	A parent holding company or control person in accordance with § 240.13d-1(b)(1)(ii)(G);
(h)	A savings associations as defined in Section 3(b) of the Federal Deposit Insurance Act (12 U.S.C. 1813);
(i)	A church plan that is excluded from the definition of an investment company under section 3(c)(14) of the Investment Company Act of 1940 (15 U.S.C. 80a-3);
(j)	A non-U.S. institution in accordance with § 240.13d-1(b)(1)(ii)(J). If filing as a non-U.S. institution in accordance with § 240.13d-1(b)(1)(ii)(J), please specify the type of institution:
(k)	Group, in accordance with Rule 240.13d-1(b)(1)(ii)(K).
Item 4.	Ownership
(a)	Amount beneficially owned: 4,200,950
(b)	Percent of class: 5.07 %
(c)	Number of shares as to which the person has:
	(i) Sole power to vote or to direct the vote: Aberdeen Group plc - 0 abrdn Inc. - 0
	(ii) Shared power to vote or to direct the vote: Aberdeen Group plc - 4,200,950 abrdn Inc. - 4,200,950
	(iii) Sole power to dispose or to direct the disposition of: Aberdeen Group plc - 0 abrdn Inc. - 0
	(iv) Shared power to dispose or to direct the disposition of: Aberdeen Group plc - 4,200,950 abrdn Inc. - 4,200,950
Item 5.	Ownership of 5 Percent or Less of a Class.
Item 6.	Ownership of more than 5 Percent on Behalf of Another Person.
	Not Applicable
Item 7.	Identification and Classification of the Subsidiary Which Acquired the Security Being Reported on by the Parent Holding Company or Control Person.
	If a parent holding company has filed this schedule, pursuant to Rule 13d-1(b)(ii)(G), so indicate under Item 3(g) and attach an exhibit stating the identity and the Item 3 classification of the relevant subsidiary. If a parent holding company has filed this schedule pursuant to Rule 13d-1(c) or Rule 13d-1(d), attach an exhibit stating the identification of the relevant subsidiary. See Exhibit
Item 8.	Identification and Classification of Members of the Group.
	Not Applicable
Item 9.	Notice of Dissolution of Group.
	Not Applicable

Item 10.

Certifications:

By signing below I certify that, to the best of my knowledge and belief, the securities referred to above were acquired and are held in the ordinary course of business and were not acquired and are not held for the purpose of or with the effect of changing or influencing the control of the issuer of the securities and were not acquired and are not held in connection with or as a participant in any transaction having that purpose or effect, other than activities solely in connection with a nomination under ?? 240.14a-11.

SIGNATURE

After reasonable inquiry and to the best of my knowledge and belief, I certify that the information set forth in this statement is true, complete and correct.

Aberdeen Group plc

Signature:	Jenny Clarke
Name/Title:	Major Shareholding Reporting Analyst
Date:	07/30/2025

abrdn Inc.

Signature:	Jenny Clarke
Name/Title:	Major Shareholding Reporting Analyst
Date:	07/30/2025

Exhibit Information

Aberdeen Group plc abrdn Holdings Limited abrdn Inc. Aberdeen Group plc is the parent company. abrdn Holdings Limited is the intermediate holding company for abrdn Inc. abrdn Inc. beneficially owns on behalf of our underlying clients 5% or greater of the outstanding shares of the security class being reported on this Schedule 13G.

FAQ

How many Codexis (CDXS) shares does abrdn Inc. own?

4,200,950 shares, representing 5.07 % of outstanding common stock.

Is the abrdn stake in Codexis considered active or passive?

The Schedule 13G was filed under Rule 13d-1(b), indicating a passive investment with no intent to influence control.

Does abrdn Inc. have sole voting rights over its Codexis shares?

No. The filing lists 0 shares with sole voting power; all 4.2 M shares are held with shared voting and dispositive power.

What type of reporting persons are Aberdeen Group plc and abrdn Inc.?

Aberdeen Group plc is a parent holding company/control person (HC); abrdn Inc. is a corporation and registered investment adviser (CO, IA).

When was the ownership position effective?

The date of event triggering the filing is 06 / 30 / 2025.