Precigen COO sells 42,703 shares in tax-related trade

Rhea-AI Filing Summary

PRECIGEN, INC. Chief Operating Officer Rutul R. Shah reported a tax-related stock sale. On May 28, 2026, he sold 42,703 shares of common stock at $4.36 per share in a sell-to-cover transaction to satisfy tax withholding obligations from restricted stock units that vested on May 23, 2026. Following this transaction, he continues to hold 497,751 shares of Precigen common stock directly, indicating he retains a substantial equity stake in the company.

Insider Trade Summary

Net Seller: 42,703 shares

Net Sell 1 txn

Insider Shah Rutul R

Role Chief Operating Officer

Sold 42,703 shs ($186K)

Type	Security	Shares	Price	Value
Sale	Common Stock	42,703	$4.36	$186K

Holdings After Transaction: Common Stock — 497,751 shares (Direct)

Footnotes (1)

1. F1. Represents the number of shares of Precigen common stock sold pursuant to a sell-to-cover transaction to satisfy tax withholding obligations upon the vesting of restricted stock units on May 23, 2026.

Key Figures

Shares sold: 42,703 shares Sale price: $4.36 per share Shares held after: 497,751 shares +1 more

4 metrics

Shares sold 42,703 shares Common stock sold on May 28, 2026

Sale price $4.36 per share Open-market sale of Precigen common stock

Shares held after 497,751 shares Direct ownership following the May 28, 2026 transaction

RSU vesting date May 23, 2026 Restricted stock units that triggered tax withholding

Key Terms

sell-to-cover, restricted stock units, tax withholding obligations, Form 4

4 terms

sell-to-cover financial

"sold pursuant to a sell-to-cover transaction to satisfy tax withholding obligations"

Sell-to-cover is when part of newly issued or exercised company stock is immediately sold to pay required taxes and fees, so the recipient keeps the remaining shares. For investors this matters because it reduces the number of shares insiders or employees actually hold after a grant, can create small, routine share sales that aren’t signal of cashing out, and slightly increases share supply on the market—like selling a portion of a paycheck to cover the tax bill.

restricted stock units financial

"tax withholding obligations upon the vesting of restricted stock units on May 23, 2026"

Restricted stock units are a type of company reward where employees are promised shares of stock, but they only fully own these shares after meeting certain conditions, like staying with the company for a set time. They matter because they can become valuable assets and are often used to motivate employees to help the company succeed.

tax withholding obligations financial

"sell-to-cover transaction to satisfy tax withholding obligations upon the vesting"

Form 4 regulatory

"INSIDER FILING DATA (Form 4)"

Form 4 is a official document that company insiders, such as executives or major shareholders, file with regulators whenever they buy or sell company shares. It provides transparency about how those with inside knowledge are trading, helping investors see if insiders are confident in the company's prospects or may be selling for personal reasons. This information can influence investor decisions by revealing insiders' perspectives on the company's value.

FAQ

AI-generated questions and answers. How Rhea-AI works. Not financial advice.

What insider transaction did PRECIGEN (PGEN) report for Rutul R. Shah?

Precigen reported that COO Rutul R. Shah sold 42,703 shares of common stock. The transaction occurred on May 28, 2026, and was described as a sell-to-cover sale to satisfy tax withholding obligations tied to recently vested restricted stock units.

At what price did the PRECIGEN (PGEN) COO sell shares in this Form 4?

The shares were sold at an average price of $4.36 per share. This open-market transaction involved 42,703 shares of Precigen common stock and was executed as part of a sell-to-cover process for tax withholding on vested restricted stock units.

Why did PRECIGEN (PGEN) COO Rutul R. Shah sell 42,703 shares?

The sale was made to cover tax withholding obligations on vested restricted stock units. The footnote explains it was a sell-to-cover transaction following the vesting of restricted stock units on May 23, 2026, rather than a discretionary reduction of his overall investment.

How many PRECIGEN (PGEN) shares does the COO hold after this transaction?

After the transaction, COO Rutul R. Shah directly holds 497,751 shares. This post-transaction ownership figure shows that despite selling 42,703 shares for tax purposes, he maintains a significant remaining position in Precigen common stock.

What type of security was involved in this PRECIGEN (PGEN) Form 4 filing?

The transaction involved Precigen common stock. The Form 4 shows a sale of 42,703 common shares at $4.36 per share in a sell-to-cover transaction related to tax withholding on vested restricted stock units.

When did the restricted stock units vest for the PRECIGEN (PGEN) COO?

The restricted stock units vested on May 23, 2026. The subsequent sale of 42,703 common shares on May 28, 2026, was specifically to satisfy tax withholding obligations arising from that vesting event, according to the footnote disclosure.

AI-generated analysis. How Rhea-AI works. Not financial advice.

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Accepted by SEC EDGAR 06/01/2026 - 04:27 PM EDT Learn about SEC filing dates

SEC Form 4

FORM 4

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934
or Section 30(h) of the Investment Company Act of 1940

OMB APPROVAL

OMB Number:			3235-0287
Estimated average burden
hours per response:			0.5

Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).

Check this box to indicate that a transaction was made pursuant to a contract, instruction or written plan for the purchase or sale of equity securities of the issuer that is intended to satisfy the affirmative defense conditions of Rule 10b5-1(c). See Instruction 10.

1. Name and Address of Reporting Person^*

Shah Rutul R

(Last) (First) (Middle)

20374 SENECA MEADOWS PARKWAY

(Street)

GERMANTOWN MARYLAND 20876

(City) (State) (Zip)

UNITED STATES

(Country)

2. Issuer Name and Ticker or Trading Symbol
PRECIGEN, INC. [ PGEN ]

5. Relationship of Reporting Person(s) to Issuer
(Check all applicable)

	Director		10% Owner
X	Officer (give title below)		Other (specify below)
	Chief Operating Officer

2a. Foreign Trading Symbol

3. Date of Earliest Transaction (Month/Day/Year)
05/28/2026

6. Individual or Joint/Group Filing (Check Applicable Line)

X	Form filed by One Reporting Person
	Form filed by More than One Reporting Person

4. If Amendment, Date of Original Filed (Month/Day/Year)

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1. Title of Security (Instr. 3)	2. Transaction Date (Month/Day/Year)	2A. Deemed Execution Date, if any (Month/Day/Year)	3. Transaction Code (Instr. 8)		4. Securities Acquired (A) or Disposed Of (D) (Instr. 3, 4 and 5)			5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4)	6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	7. Nature of Indirect Beneficial Ownership (Instr. 4)
			Code	V	Amount	(A) or (D)	Price
Common Stock	05/28/2026		S		42,703(1)	D	$4.36	497,751	D

Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned (e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security (Instr. 3)	2. Conversion or Exercise Price of Derivative Security	3. Transaction Date (Month/Day/Year)	3A. Deemed Execution Date, if any (Month/Day/Year)	4. Transaction Code (Instr. 8)		5. Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5)		6. Date Exercisable and Expiration Date (Month/Day/Year)		7. Title and Amount of Securities Underlying Derivative Security (Instr. 3 and 4)		8. Price of Derivative Security (Instr. 5)	9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4)	10. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	11. Nature of Indirect Beneficial Ownership (Instr. 4)
				Code	V	(A)	(D)	Date Exercisable	Expiration Date	Title	Amount or Number of Shares

Explanation of Responses:

1. Represents the number of shares of Precigen common stock sold pursuant to a sell-to-cover transaction to satisfy tax withholding obligations upon the vesting of restricted stock units on May 23, 2026.

	/s/ Rutul R. Shah, by Donald P. Lehr, as attorney-in-fact	06/01/2026
	** Signature of Reporting Person	Date
Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
* If the form is filed by more than one reporting person, see Instruction 4 (b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
Note: File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB Number.
* Form 4: SEC 1474 (03-26)