Nautilus director receives 45,000 stock options

Nautilus Biotechnology director Melissa B. Epperly received a grant of stock options covering 45,000 shares of common stock.

Rhea-AI Filing Summary

Nautilus Biotechnology director Melissa B. Epperly received a grant of stock options covering 45,000 shares of common stock. The options have an exercise price of $2.28 per share and expire on June 18, 2036.

These options vest in equal monthly installments over 12 months, with one-twelfth of the grant vesting each month after the grant date, as long as she continues to serve as a “Service Provider” under the company’s 2021 Equity Incentive Plan. Following this grant, she holds 45,000 stock options directly.

Insider Trade Summary

Grant/Award: 45,000 shares

Grant/Award 1 txn

Insider Epperly Melissa B,

Role Director

Type	Security	Shares	Price	Value
Grant/Award	Stock Option (Right to Buy)	45,000	$0.00	$0.00

Holdings After Transaction: Stock Option (Right to Buy) — 45,000 contracts (Direct)

Footnotes (1)

1. F1. Subject to reporting person's continuous status as a "Service Provider" (as defined in the the Issuer's 2021 Equity Incentive Plan) through each applicable vesting date, one-twelfth (1/12th) of the shares subject to the Option shall vest on a monthly basis following the date of grant on the same day of the month as such grant date (or on the last day of the month, if there is no corresponding day in such month).

Key Figures

Options granted: 45,000 options Exercise price: $2.28 per share Expiration date: June 18, 2036 +3 more

6 metrics

Options granted 45,000 options Stock Option (Right to Buy) grant to director

Exercise price $2.28 per share Conversion/exercise price of granted options

Expiration date June 18, 2036 Option term end date

Underlying shares 45,000 shares Common stock underlying the options

Post-grant option holdings 45,000 options Total derivative holdings following transaction

Vesting schedule 1/12 monthly One-twelfth of options vest each month after grant

Key Terms

Stock Option (Right to Buy), 2021 Equity Incentive Plan, Service Provider, vesting

4 terms

Stock Option (Right to Buy) financial

"security_title: Stock Option (Right to Buy)"

2021 Equity Incentive Plan financial

"defined in the the Issuer's 2021 Equity Incentive Plan"

Service Provider financial

"continuous status as a "Service Provider""

vesting financial

"shares subject to the Option shall vest on a monthly basis"

Vesting is the process by which you earn full ownership of something, like company stock or a retirement benefit, over time. It’s like earning the right to keep a gift piece by piece the longer you stay with a company, making sure employees stay committed before they receive all the benefits.

FAQ

AI-generated questions and answers. How Rhea-AI works. Not financial advice.

What did Melissa B. Epperly receive in this Nautilus (NAUT) Form 4 filing?

Melissa B. Epperly received a grant of 45,000 stock options for Nautilus Biotechnology common stock. These options are a form of compensation that gives her the right to buy shares in the future at a fixed exercise price of $2.28 per share.

What is the exercise price and expiration date of Epperly’s Nautilus (NAUT) options?

The granted options have an exercise price of $2.28 per share and an expiration date of June 18, 2036. This means she can choose to exercise the options at $2.28 any time before they expire, subject to vesting.

How do the 45,000 Nautilus (NAUT) options granted to Epperly vest?

The 45,000 options vest in monthly installments. One-twelfth of the options vest each month after the grant date, on the same calendar day, provided she continuously qualifies as a “Service Provider” under Nautilus Biotechnology’s 2021 Equity Incentive Plan.

What ongoing service conditions apply to Epperly’s Nautilus (NAUT) stock options?

The options vest only if she maintains continuous status as a “Service Provider” under the Nautilus Biotechnology 2021 Equity Incentive Plan. If that status ends, any unvested portion would stop vesting and could ultimately be forfeited under plan terms.

How many Nautilus (NAUT) options does Epperly hold after this Form 4 transaction?

After this grant, Melissa B. Epperly holds 45,000 stock options directly. These options each relate to one share of Nautilus Biotechnology common stock and together represent her current reported derivative position from this particular grant transaction.

AI-generated analysis. How Rhea-AI works. Not financial advice.

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Accepted by SEC EDGAR 06/22/2026 - 04:13 PM EDT Learn about SEC filing dates

SEC Form 4

FORM 4

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934
or Section 30(h) of the Investment Company Act of 1940

OMB APPROVAL

OMB Number:			3235-0287
Estimated average burden
hours per response:			0.5

Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).

Check this box to indicate that a transaction was made pursuant to a contract, instruction or written plan for the purchase or sale of equity securities of the issuer that is intended to satisfy the affirmative defense conditions of Rule 10b5-1(c). See Instruction 10.

1. Name and Address of Reporting Person^*

Epperly Melissa B,

(Last) (First) (Middle)

C/O NAUTILUS BIOTECHNOLOGY, INC.

2701 EASTLAKE AVE. E.

(Street)

SEATTLE WASHINGTON 98102

(City) (State) (Zip)

UNITED STATES

(Country)

2. Issuer Name and Ticker or Trading Symbol
Nautilus Biotechnology, Inc. [ NAUT ]

5. Relationship of Reporting Person(s) to Issuer
(Check all applicable)

X	Director		10% Owner
	Officer (give title below)		Other (specify below)

2a. Foreign Trading Symbol

3. Date of Earliest Transaction (Month/Day/Year)
06/18/2026

6. Individual or Joint/Group Filing (Check Applicable Line)

X	Form filed by One Reporting Person
	Form filed by More than One Reporting Person

4. If Amendment, Date of Original Filed (Month/Day/Year)

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1. Title of Security (Instr. 3)	2. Transaction Date (Month/Day/Year)	2A. Deemed Execution Date, if any (Month/Day/Year)	3. Transaction Code (Instr. 8)		4. Securities Acquired (A) or Disposed Of (D) (Instr. 3, 4 and 5)			5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4)	6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	7. Nature of Indirect Beneficial Ownership (Instr. 4)
			Code	V	Amount	(A) or (D)	Price

Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned (e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security (Instr. 3)	2. Conversion or Exercise Price of Derivative Security	3. Transaction Date (Month/Day/Year)	3A. Deemed Execution Date, if any (Month/Day/Year)	4. Transaction Code (Instr. 8)		5. Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5)		6. Date Exercisable and Expiration Date (Month/Day/Year)		7. Title and Amount of Securities Underlying Derivative Security (Instr. 3 and 4)		8. Price of Derivative Security (Instr. 5)	9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4)	10. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	11. Nature of Indirect Beneficial Ownership (Instr. 4)
				Code	V	(A)	(D)	Date Exercisable	Expiration Date	Title	Amount or Number of Shares
Stock Option (Right to Buy)	$2.28	06/18/2026		A		45,000		(1)	06/18/2036	Common Stock	45,000	$0	45,000	D

Explanation of Responses:

1. Subject to reporting person's continuous status as a "Service Provider" (as defined in the the Issuer's 2021 Equity Incentive Plan) through each applicable vesting date, one-twelfth (1/12th) of the shares subject to the Option shall vest on a monthly basis following the date of grant on the same day of the month as such grant date (or on the last day of the month, if there is no corresponding day in such month).

Remarks:

	/s/ Matthew B. Murphy, as Attorney-in-Fact	06/22/2026
	** Signature of Reporting Person	Date
Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
* If the form is filed by more than one reporting person, see Instruction 4 (b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
Note: File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB Number.
* Form 4: SEC 1474 (03-26)